of Fat in the Alimentary Canal. 



255 



were lost in the process of separation. We, however, see that only 

 0*402 gram of the total ether extract was left in the large intestine, 

 so that 8*457 grams of the fat had really been absorbed during the 

 seven hours' digestion ; that is to say, as much as 85*98 per cent, 

 of the total quantity of fat that had traversed the pylorus. 



Table III. — Normal Dog, weighed 10*35 kilos., killed by Chloroform 

 Seven Hours after feeding with Milk. 





Total ether 

 extract. 



Neutral fat. 



Free fat 

 acids. 



Fat acids as 

 soaps. 





Total. 



p.c. 



Total. 



p. c. 



Total. 



P- 



c. 



Total. 



p. c. 





12 -027 



100 



11 -533 



95*89 



0'455 



3 



78 



0-039 



0*33 



Found in — 























8-116 



100 



6*061 



74-67 



2-012 



24 



81 



0-043 



0-52 



Small intestine . 



0*623 



100 



0-080 



12-84 



0-522 



83 



46 



0-021 



3-70 



Large intestine. 



0-741 



100 



0-229 



30-91 



0-452 



61 



•03 



0-059 



8-06 



In Table III the total fat given was 12*027 grams, and after seven 

 hours' digestion 8*116 grams remained in the stomach. The stomach 

 had consequently discharged 3*911 grams of the fat into the intes- 

 tines (i.e., 32*52 per cent.). 



The fat that remained in the stomach during the seven hours had 

 undergone similar changes to those already described in the former 

 two experiments. The neutral fat had decreased from 11*533 grams 

 in the milk given to 6*061 grams. Thus, while the milk fat contained 

 95*89 per cent, as neutral fat, the stomach only contained 74 - 67 per 

 cent., so that at least 21*22 per cent, of the neutral fat given had been 

 split up in the stomach into free fat acids or traversed the 

 pylorus. 



The free fat acids on the other hand had increased very markedly ; 

 for whereas the milk contained but 0'455 gram the stomach con- 

 tained 2*012 grams, the percentage composition having thus changed 

 from 3*78 to 24 81. 



The soap had also increased in quantity in the stomach from 

 0*039 gram, in the milk given, to 0*043 gram, which altered the 

 percentage composition from 0*33 to 0*52 per cent. 



Thus, in aJl three experiments we have confirmation of the fact 

 that neutral fat is split up into free fat acids while in the stomach 

 itself, and that some of these free fat acids formed soaps. 



With regard to the small intestines we find only 0'623 gram of 

 ether extract was present, while we know that 3'91 grams had 



