No. 566] INHERITANCE IN EARS OF MAIZE 107 



latter, owing to the rarity of 8 in male gametes. Only 7 

 such red ears were tested and all yielded red and varie- 

 gated ears in typical Mendelian ratios, showing that all 

 of them were VS like any F x hybrid between red and 

 variegated races. Of two F 2 reds from self ed F 1 % one 

 again yielded reds and variegates and one apparently 

 bred true red. Three F 2 reds, from F x reds crossed by 

 whites, yielded reds and whites only — typical Mendelian 

 results throughout. 



When F x red-eared plants arose from either homozy- 

 gous or heterozygous, variegated ears that had been cross- 

 pollinated by whites they yielded only red-eared and 

 white-eared, never variegated-eared, offspring (see page 

 100), just as if they were F 1 ears of a cross of reds with 

 whites. By hypothesis the parent variegated-eared plants 

 were V — and VV, and their red grains S— and SV (or 

 possibly 88). The gametes associated with such grains 

 were therefore 8 and — , and 8 and V (or possibly all S). 

 The male gametes from white races were all — . The F, 



plants were therefore 8— t V—, and , only those with 



8 — having red ears. The five red-eared F t plants that 

 were tested produced in F 2 red-eared and white-eared 

 plants in Mendelian ratios. Of the F 2 red-eared plants 

 one bred true in F 3 and three again segregated into reds 

 and whites. 



When heterozygous, variegated, parent ears were self- 

 pollinated, the Fj red-eared plants behaved in some cases 

 like hybrids of red with variegated races and in other 

 cases like hybrids of red with white races (see page 99). 

 Our assumption is that the variegated-eared parent plants 

 were V — and their red grains 8—. The gametes asso- 

 ciated with these red grains were of course S and — . The 

 male gametes of the same plants were doubtless largely 

 V and — , though a few were probably 8. The F x plants 



must therefore have been , V — , S— SV or SS. Beds 



with 88 would be expected only rarely, and of the 11 F 1 

 reds tested none had that formula, else they would have 

 bred true in F 2 . Seven of the 11 F x reds evidently were 



