No. 577] SHORTER ARTICLES AND DISCUSSION 47 



it falls off toward the stigmatic end, where the fecundity 

 may be even lower than it is a little farther down in 

 the pod. This is admirably shown in Diagram 3, in which GO 

 stands for a series of Burpee 's Stringless grown at the Missouri 

 Botanical Garden, NH for a series of Navy beans grown at 

 Sharpsburg, Ohio, and LL for a series of Golden Wax beans 

 grown at Lawrence, Kansas. All were grown in 1907. Here the 

 percentage of development of ovules at different positions in the 

 pod is shown for the different classes of pods by the scales to the 

 left of the figures. The reader may ascertain the class of pods 

 represented by any particular curve by noting the number of 

 circles representing percentage development in the various posi- 

 tions. These correspond to the number of ovules per pod. In the 

 diagrams the positions (abscissae) from left to right represent the 

 positions from the base to the tip of the pod. 



J. Arthur Harris 



ANOTHER GENE IN THE FOURTH CHROMOSOME OP 

 DROSOPHILA 



Until the appearance of bent wings, only three groups of 

 linked genes had been found in Drosophila amelophila, although 

 four pairs of chromosomes had been identified in the diploid 

 group. Since the character bent wings, worked out by Mr. H. J. 

 Muller, was found to be unassociated with any of the three 

 groups, the gene producing this character was said to be located 

 in the fourth chromosome. 



Recently a new character, designated as eyeless, appeared. 

 Flies having this character either lacked eye pigment and om- 

 matidia or had one or both eyes reduced in size. All of the pure 

 stock showed some loss of eye structures. Eyeless is recessive to 

 the normal eye. In order to determine the linkage, eyeless males 

 were crossed in turn to females of the stocks at Columbia Uni- 

 versity. These stocks representing the three groups were (1) 

 miniature wings, (2) black body and vestigial wings, and (3) 

 spread wings. The genes producing these characters are in the 

 first, second and third chromosomes, respectively. The F 1 from 

 all three crosses had normal eyes. They were inbred in each 

 case and gave the following. 



The equation should be w = 9.987 + .021 p. The line as it appears here is 

 correctly drawn. 



