560 Tin: am eric ax naturalist [Vol.l 



produced, has been found. This very evidently shows 

 that variation in the amount of suppression of the de- 

 terminer for maleness takes place in the determiner for 

 sex of these normal grains. This normal pollen when 

 used to pollinate pistillate flowers should give, in some 

 cases, females bearing reflexed stamens and in others 

 hermaphrodites, depending upon the extent to which sup- 

 pression of maleness is. lacking in the chromosome bear- 

 ing the sex determiners of these normal pollen grains. 



From the cross "R xU" (female FF X hermaphrodite 

 HF) should be expected females (FF) and hermaphro- 

 dites (HF) in the proportion of 1:1. This ratio is met 

 exactly in the cross "Rx"0"" = 207 upright and 206 

 reflexed. 



The following analysis, kindly furnished me by Mr. 

 Anthony of the New York State Agricultural Experiment 

 Station, of the cross "hermaphrodite female X pure 

 male" which produced "56 hermaphrodites + 51 males," 

 shows that both hermaphrodites and females were used as 

 the female parent and that three kinds of males were 

 used, namely, wild males, males one generation from the 

 wild and "intermediates" (males bearing occasionally a 

 few well developed pistils). 



The various combinations will be considered separately. 



The cross hermaphrodite (FH) X wild male (FM) pro- 

 duced 7 hermaphrodites, 6 females and 9 males, somewhat 

 approximating the expected ratio of 1 female (FF) : 1 



5 The result of hermaphrodite X wild male. 



6 These totals do not quite coincide with those given in the published data, 

 as the parents of 7 of the vines were not certainly known and are therefore 



