562 THE AMEBIC AX NATURALIST [ Vol. L 



Farm showed that certain clusters of a vine may be 

 entirely staminate, while others of the same vine con- 

 tain all gradations from staminate to functionally per- 

 fect flowers, many of which are capable of setting fruit. 

 There is very evidently a suppression of femaleness in 

 certain parts of these plants and not in others. This 

 raises the question as to whether pollen from the pure 

 staminate clusters can transmit only determiners for 

 maleness and femaleness, or whether they are able to 

 transmit the hermaphrodite condition. Mr. Anthony in- 

 forms me that the pollen used in the above cross was 

 "most certain to have come from such blossoms" (i. e., 

 from pure male clusters). If the two types of gametes 

 produced by these flowers bear the determiners H and F. 

 respectively, the cross hermaphrodite X intermediate 

 should produce hermaphrodites and females in a 3:1 

 ratio and no males, while if these male flowers function 

 as normal males and the gametes produced carry the 

 determiners F and M respectivelv, a ratio of 1 hermaph- 

 rodite (HF) : 1 female (FF) : *2 males (HM and FM) 

 would be expected. A close approximation to this ratio 

 was actually produced. 



The cross female (FF) X intermediate which produced 

 1 female and 3 males, gives further evidence that the 

 staminate flowers of the intermediate vines do not pro- 

 duce gametes bearing the hermaphroditic determiner, but 

 act as pure males. Otherwise the appearance of the 

 three males can not be explained. 7 Although the number 

 of vines produced from this cross is small, still the appear- 

 ance of the three males is extremely significant. 



It has already been pointed out that in the wild there 

 are two types of vines, male and female, and that under 

 cultivation a third type, the functional hermaphrodite, is 

 common. We are now in a position to discuss the pos- 

 sible origin of these types. It is clear that both the 

 staminate and functionally pistillate vines carry the de- 

 terminers for femaleness and maleness, respectively, 



'Anthony (1914) pointed out the fact that the pollen from these inter- 



