No. 597] INHERITANCE OF SEX IN THE GRAPE 563 



partially suppressed and therefore, there are two possi- 

 bilities with regard to the origin of functional hermaph- 

 rodites. (1) Maleness may express itself fully in one 

 of the chromosomes bearing the determiner for female- 

 ness in a pistillate plant, and (2) femaleness may express 

 itself fully in the chromosome bearing the male de- 

 terminer in the staminate plant. I think it oan be said 

 definitely that functional hermaphrodites have been de- 

 veloped in both of these ways. The production of her- 

 maphrodites from the cross female X female can hardly 

 be explained on any other basis than entire lack of sup- 

 pression of maleness in certain gametes bearing the tenia I e 

 determiner, while the appearance of well-developed pistils 

 in a few flowers of certain male vines must be the result 

 of lack of suppression of femaleness in at least a portion 

 of the somatic cells of these males. 



As there is an apparent segregation in the somatic 

 tissue of these vines, whole clusters and occasionally all 

 clusters on a cane being staminate while others bear 

 many intermediate and perf ect flowers, it seems logical to 

 assume that the perfect flowers can transmit the her- 

 maphroditic condition to some of their seedlings through 

 both the male and the female gametes, resulting in either 

 homozygous or heterozygous hermaphrodites, all of whose 

 flowers are perfect. 



LITERATURE CITED 



