Milou Deer 233 



deposits of Greece and France. In the latter kind the main fork of the 

 antler is placed still lower down, so that the undivided portion of the beam 

 is less than a third the total length of the antler ; the front prong of the 

 main fork being relatively longer. I can find no sufficient characters for 

 regarding C. neschersensis, which occurs in the same beds, as distinct from 

 the present form. 



Distribution. — France during the later part of the Pliocene period, the 

 typical localities being Perrier and Arde in the Department of Puy-de-D6me. 



X. The Milou Deer — Genus elaphurus 



E/ap/iurus, Milne-Edwards, Ann. Sci. Nat. ser. 5, vol. x. p. 380 (1866) ; 

 Gray, Cat. Ruminants Brit. Mus. p. 82 (1872) ; Fitzinger, SB. Ak. Wien, vol. 

 lxx. part i. p. 328 (1874) ; Brooke, Proc. Zool. Soc. 1878, p. 906, as a sub- 

 genus of Cervus. 



Characters. — Lateral metacarpal bones as in Cervus, that is to say, 

 represented by their upper extremities. Antlers large and cylindrical, with 

 the beam making a marked angle with the plane of the face ; no brow-tine 

 or sub-basal snag, the beam dichotomously forking at a comparatively small 

 distance above the burr ; the front, or upper prong of the main fork curving 

 forwards and again dividing once or more, the posterior, or lower prong long, 

 straight, simple, and projected backwards. Muzzle with a large naked 

 portion, which is deeper and broader below the nostrils than in the elaphine 

 group, but extends only a little on to the front of the face, where its upper 

 border is deeply concave ; ears small and narrow ; tail very long, cylindrical, 

 and bushy at the extremity ; neck maned in the male ; face long. Pelage 

 of adult uniformly coloured throughout the upper-parts ; in young spotted. 

 No tarsal tuft. Metatarsal tuft continuous and situated in the upper third 

 of the metatarsus. The gland-pit in the skull large, and the face-gland also 

 large. Main hoofs large and spreading ; lateral hoofs very large. Upper 

 canines small ; upper molars moderately tall, with a small additional column 

 on the inner side. The vomer not dividing the inner aperture of the nostrils 

 on the under surface of the skull into two moieties. Size large ; build 

 heavy, with the limbs stout. No foot-glands. 



Much difference of opinion has existed as to the systematic position of 



2 H 



