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MEMOIRS OF THE NEW YORK BOTANICAL GARDKN 



[VOL. 10 



detail, cell walls of varied thickness are represented as follows: thin-walled, a 

 single line separating- cells; moderately thick-walled, a pair of lines separating 

 cells; thick-walled sclerenchyma or tracheary elements, a pair of concentric lines 

 representing the thickness of the wall for each cell. Obviously, some degree of 

 integration between these three types occurs, so the representations are not pre- 

 cise, although they are approximately correct. 



In diagrammatic drawings, patterns have been used to differentiate cell types 

 and tissues. These are as follows: stippled, sclerenchyma; white, parenchyma 

 (and epidermis, if unsclerified) ; black, xylem ; cross-hatched, phloem; hori- 

 zontal lines, thin-walled sclerenchyma or thick-walled parenchyma (walls show- 

 ing lignification) ; wide vertical lines, cork; narrow vertical lines, laticiferous 

 cells. 



In the anatomical descriptions which follow, the genera of Gochnatinae 

 (Stenopadus, Stomatochaeta, Chimantaea, and Quelchia) are considered first. 

 These are followed by genera belonging to Gerberinae (Mutisinae of Hoffmann, 

 1890), Gongylolepis and its allies (Achnopogon, Neblinaea, and Duidaea). The 

 rather isolated monotypic genus Glossarion is considered" last in each case. 



I. STEM 



Cortex and Pith. Within Gochnatinae, Stenopadus, Stomatochaeta, and Chi- 

 mantaea show a similar basic pattern. Stenopadus (fig. 1) lias a cortex contain- 

 ing numerous thick-walled sclereids, developed individually or in irregular 

 groups. Cortical leaf traces show a prominent cap of fibers which, on larger 

 traces, extend around the bundle (top of fig. 1). Fibrous bundles are also formed 

 on strands of the vascular cylinder. During secondary growth, these are aug- 

 mented by sclerification of parenchyma cells in interfascicular areas, so that 

 a nearly continuous cylinder of sclerenchyma is present around the phloem of 

 such a stem. In the pith (fig. 1, right), nests of thick-walled sclereids are em- 

 bedded in a background of thin-walled sclerenchyma. Occasional areas of thin- 

 walled parenchyma may also be present in pith of species of Sit nopadus. In the 

 details mentioned above, other species of Stenopadus examined (S. stipitatus, 

 S. cucullatus) were found to be identical. 



The relatively narrower stems of Stomatochaeta (fig. 2) conform to much 

 the same pattern. The primary stem shown is characterized by a sclerified epi- 

 dermis covered with a prominent cuticle. In the cortex, thick-walled sclereids 

 are present. Fibrous bundle caps occur, and during secondary growth a few 

 interfascicular parenchyma cells sclerose, although not enough to form the con- 

 spicuous cylinder seen in Stenopadus stems. In the pith, nests of sclereids are 

 surrounded by thin-walled parenchyma. During secondary growth, this paren- 

 chyma becomes lignified, so that mature pith is indistinguishable from that of 

 Stt nopadus. 



Chimantaea (fig. 3) shows many of the same features, seemingly modified 

 in accordance with the very different growth habit of this genus. Chimantaea 



Fig. 1. Stenopadus connellii, Wur black 34265 ; portion of transection of stem with second- 

 ary growth; periderm at left; xylem and pith at right; black areas among cells represent 

 carbonized resin deposits. Fig. 2. Stomatochaeta condensata, Maguire 40603 ; portion of tran- 

 section of primary stem, epidermis at left. Fig. 3. Chimantaea rupicola, Styermark Sr Wur- 

 dack 748; diagram of transection of stem with secondary growth, periderm at left; symbols 

 explained in text. 



