s 



56 MEMOIRS OF THE NEW YORK BOTANICAL GARDEN [VOL. 10 



less advanced than the smaller, reduced glands in the carpels of Diomma and 

 Sohnreyia. Arguments for and against the primitiveness or advancedness of 

 certain characteristics in plants can sometimes be interpreted in two ways. The 

 above, therefore, is only meant to state the authors' opinions. 



Actually the occurrence of the schopfbaumartig habit and monocarpy in the 

 three genera under discussion is based on a relatively small body of information. 

 If it is subsequently shown that these two features are in truth constant char- 

 acters of all species of Spathelia, Diomma, and Sohnreyia, it will strengthen the 

 structure of Spatlielioicleae sensu novo. Further, more detailed studies, on a 

 wide variety of material, are desirable to determine the exact nature of the 

 stamina! filament proliferations described for these genera. 



CONCLUSIONS 



1. The genera Diomma, incertae sedis, and Sohnreyia, Eutaceae, are closely 

 related. 



2. Diomma is definitely a member of Eutaceoi (sensu Engler 1931) as is 

 shown by investigations of floral morphology, pollen structure, secretory ele- 

 ments, nodal anatomy, petiole vascular structure, and wood anatomy. 



3. Diomma and Sohnreyia should be included in the subfamily Spatheli- 

 oideae, sensu novo, as they are both closely related to Spathelia. 



ACKNOWLEDGMENTS 



For their interest in suggesting the problem and their help throughout the 

 progress of the study, the authors express their gratitude to Dr. Bassett Maguire 

 and Dr. John J. Wurdack of the Xew York Botanical Garden and Dr. Richard 

 S. Cowan of the U. S. National Herbarium. The following botanists were very 

 kind in arranging for loans of study specimens from their respective institutions 

 and the authors appreciate their efforts : Dr. Theodor Just of the Chicago Nat- 

 ural History Museum, Mr. AVilliam T. Stearn of the British Museum, Dr. G. 

 Taylor of the Royal Botanic Gardens, Kew, Dr. H. J. Lam of the Rijksherbar- 

 ium, Leiden, Dr. Charles Baehni of the Conservatoire et Jardin Botaniques, 

 Geneva, and Dr. John R. Reeder of Yale University. For his kindness in exam- 

 ining pollen specimens the authors are much indebted to Dr. G. Erdtman of the 

 Palynological Laboratory in Stockholm. Dr. Richard A. Howard of the Arnold 

 Arboretum gave of his time to explain methods for studying petiole sections. 

 Professor I. AY. Bailey of Harvard University deserves especial thanks for his 

 critical reading of the manuscript and his suggestions for improvement. 



Literature Cited 



Agardh, J. G. 1858. Theoria systematis plantarum. 1-404. 

 Bentham, G. & Hooker, J. D. 1862. Genera plantarum. 1(1): 1-434. 



Blenk, P. 1884. Ueber die durchsichtigen Punkte in den Blattern. Flora 67: 49-57, 97-112, 



136-144, 204-210, 223-225, 275-283, 291-299, 339-349, 355-370, 371-386. 

 Bremekamp, C. E. B. 1944. Materials for a monograph of the Strobilanthinae (Acanihaceae). 



Verh. Nederl. Akad. Wet. Sect. 2. 41(1): 1-306. 

 Chattaway, M. Margaret. 1932. Proposed standards for numerical values used in describing 



woods. Trop. Woods 29: 20-28. 

 Chowdhury, K. A. 1939. The formation of growth rings in Indian trees. Indian For. Eec. 



II. Utilization 2(1): 1-39. 



