1960] 



TAXOXOMIC RELATIOXSHIPS OF DIOMMA 



63 



partial key and by descriptions presented by Leon & Alain (1951). One further, 

 but doubtful, species from Mexico has been proposed by Sesse and Mocino. 



Spathelia Subg. 2. Diomma (Engler ex Harms) Cowan & Brizicky, stat. nov. 



Diomma Engler ex Harms, Nat. Pflanzenfam. ed. 2. 19a: 460. 1931. 



Type Species: Spathelia (subg. Diomma) ulei (Engler ex Harms) Cowan & 

 Brizicky, comb. nov. (based on Diomma ulei Engler ex Harms, I.e.) 



Key to the Species of Subg. Diomma 



1. Leaflets with glandular punctae scattered, not limited to the margins. Fruits not 

 over 2.5 cm wide, deeply cordate basally, retuse apically, each half about semi- 

 circular, i. S. fruticosa. 



1. Leaflets with glandular punctae restricted to the margins. Fruits larger, differently 



shaped. 2. 



2. Filaments not aliforai basally or if slightly expanded, not aliform and not bidentate 



at the apex. Pedicels 4—5.5 mm long. Ovary broadest basally ; fruits broadly ovate, 

 attenuate apically, the seed-bearing portion narrowly lanceolate. 2. S. ulei. 



2. Filaments winged basally. Pedicels 1.5-2.5 mm long. Ovary broadest at about the 



middle; fruits oblate or obovate-oblate, not attenuate apically, the seed-bearing 

 portion broadly ovate. 3. 



3. Median leaflets about 10-12 cm long, oblong, acute, the rachis ca. 30-45 cm long. 



Fruits glabrous, oblate, slightly apiculate apically, 4 cm broad. 3. S. chimantaensis. 



3. Median leaflets about 14 cm long, oblong-lanceolate, acuminate, the rachis 47-57 cm 



long. Fruits puberulous, obovate-oblate, truncate apically, 3 cm broad. 4. S. neblmaensis. 



1. Spathelia fruticosa (Steyermark) Cowan & Brizicky. comb, nov. Fig. 1, a. 

 Diomma fruticosa Steyermark, Fieldiana Bot. 28: 272. 1952. 



Tree 1-5 m tall, the tip of the stem densely puberulous minutely. Leaves 

 25-30-foliolate, the leaflets with scattered glandular punctae, the petioles terete 

 except at the canaliculate base, 10-18.5 cm long, densely puberulous but glabres- 

 cent, the rachis 42-54 cm long, densely puberulous, glabrescent, the leaflets op- 

 posite to alternate, the petiolules 1-2.5 mm long, puberulous but glabrescent, the 

 blades glabrous at maturity, those nearest the leaf base smaller, ca. 9-10 cm long, 

 3.5-4 cm wide, oblong-ovate, the blades of the upper leaflets larger, slightly to 

 strongly arcuate, oblong-lanceolate or oblong. 9-15 cm long, 3-5 cm wide, the base 

 of all the leaflets inequilateral, the upper side rounded-obtuse, the lower side 

 subobtuse (or rarely acute) to rounded-obtuse, the apex abruptly acute to 

 acuminate with the tip blunt, the venation plane on the upper leaflet surface or 

 the costa slightly salient, the eosta and the primary vein-branches salient beneath. 

 Inflorescences 35-63 cm long, the axis densely puberulous, the bracts about 1 mm 

 long, puberulous, the pedicels 1-2 mm long, puberulous. Sepals 1.5-2 mm long, 

 0.8-1.2 mm wide, oval to narrowly elliptic, emarginate apically. glabrous ex- 

 cepting the ciliolate margin; petals 3.5-4 mm long, 2 mm wide, oval, glabrous. 



Fig. 1. a, Spathelia fruticosa, fruit, X 1 (drawn from SteyermarTc 60327). b-d, S. ulei. 

 b, fruit, X 1; c, filament, X 7.5; d, ovary, X 10 (drawn from Vie 8646). e-f, S. neblinaensis. 

 e, leaflet, X 0.5; /, immature (?) fruit, X 1 (drawn from Maguire, WurdacTc 4' Ifaguire 

 42329). g-4, S. chimantaensis. g, fruit, X. 1; h, filament, X 7.5; i, leaflet, X 0.5 (drawn from 

 SteyermarTc 75699). j-m, S. excelsa. j, fruit, X 1; Tc, lateral view of filament, X 7.5; I, view 

 of outer face of filament, X 7.5; m, gynoecium, X 7.5 (j drawn from Ducke 20472, Tc-m drawn 

 from Ducke 15702). 



