1961] 



BOTANY OF THE GUAYANA HIGHLANDS PART IV (2) 



19 



Pentaspatella ramosa Gleason. 



Sauvagesia duclcei Sleumer, Eepert. Sp. Nov. 42: 263. 1937. 



Dwyer (in Fordham University library, thesis) had proposed the transfer of 

 P. ramosa to Leitgebia; both L. guianensis and P. ramosa have the same small 

 inner staminodia and placentae limited to the ovary base. We feel however that 

 the vast habital difference warrants the generic retention of Pentaspatella; it 

 probably represents a distinct divergence from Sauvagesia, with the final floral 

 reduction the same as in Leitgebia. We have examined Venezuelan material 

 of Pentaspatella from the Yapacana savannas and Esmeralda Sabana Grande, 

 as well as both topotypic (Ducke 999) and isotypic (Ducke H.J.B.R. 30124, US) 

 material of 8. duckei and have noted no differences between the Venezuelan and 

 Brazilian collections. Dwyer 's placentation character for Sauvagesia is certainly 

 valid within the limits of our generic sampling; S. erecta L., S. imthurniana 

 (Oliver) Dwyer, S. sprengelii St.-HiL, 8. nudicaulis Maguire & Wurdack, and 

 S. smithiana Dwyer all have distinct ovary wall placentae. 



Adenanthe Maguire, Steyermark & Wurdack, gen. nov. 



Inflorescentia paniculata ; bracteis dissectis glandulosis; sepalis 5, quincun- 

 e'ialibus; androgynophoro androeciali cupuliformi ; staminodiis 1-seriatis; stamin- 

 ibus 5; pistillo 2-carpellato, placentis parietalibus ; capsula bivalvata; semi- 

 nibus elongata alatis. Frutex; foliis alternatis; stipulis similibus aequalibus 

 ovatis. 



Typus: Adenanthe bica/rpellata Maguire, Steyermark & Wurdack. 



The name is from aden, aS^v, gland, and anthe, <zv0y;, flower. 

 Adenanthe bicarpellata Maguire, Steyermark & Wurdack, sp. nov. Fig. 27 A, B. 



Frutex - pauciramosus 0.3-3.0 m altus; ramulis 8-10 mm diam.; foliis sessi- 

 libus saepe dense confertis, laminis ellipticis vel oblanceolatis, coriaceis, 3-11 cm 

 longis, 1.5-4.0 cm latis, ad apicem obtusis vel acutiusculis, venis numerosis pro- 

 minulis ad plus minusve 80° angulos adscendentibus ; stipulis aequalibus ovatis 

 vel lanceolatis, 1.0-1.5 cm longis, 5-7 mm latis, ovatis vel lanceolatis, acutis, 

 caducis; inflorescentiis terminalibus paniculatis, 15-30 cm longis, ramulis 

 lateralibus brevissimis vel 12-15 cm longis, bracteolis dissectis, glandulosis; 

 sepalis 5, quincuncialibus ovato-lanceolatis vel lanceolatis acutis 3-5 mm latis, 

 8-13 mm longis, inconspicue 9-11-parallelivenatis, marginibus valde pectinate 

 stipitato-glandulosis, stipitibus 2.5-3.5 mm longis; petalis 8-12 mm latis, 18-22 

 mm longis, obovatis obtusis, inconspicue punctato-glandulosis ; staminodiis 10, 

 1-seriatis, in corona ca. 2 mm alta dispositis, staminodiis staminibus alternatis 5, 

 anguste spathulati-oblongis, obtusis, interne ad basim carinatis, 8-9 mm longis, 

 1.1-1.3 mm latis,- staminodiis staminibus oppositis 5, anguste lanceolatis anguste 

 acutis, 9-10 mm longis, 0.5-0.8 mm latis; staminibus 5, subsessilibus introrse lin- 

 eari-laneeolatis acutis 5-6 mm longis, ca. 1.2 mm latis, 4-thecatis, thecis posteriori- 

 bus 0.2-0.3 mm productis, lateraliter dehiscentibus a poris elongatis subapicali- 

 bus; pistillis 2-carpellatis, 1-locularibus placentis parietalibus 2, ovulis numerosis; 

 stylo subulato ca. 7-8 mm long; capsula ovata acuta, subcompressa, submem- 

 branacea, bivalvata, 8-10 mm alta, suturis non intrusis; seminibus 3.0-3.5 mm 

 longis, plus minusve oblongis alatis alveolatis. 



VENEZUELA: Bolivar: Chimanta Massif, central section, abundant and dominant 

 around margins of savannas, swampy savanna above Summit Camp, alt. 1940 m; petals smaller 

 than in 375-A, elliptic-oblong to ovate-oblongate, white with pink tips, inflorescence and sepals 

 wine-red; 3 Feb 1955, Steyermarlc Wurdack 377 (holotype, NY). Chimanta-tepm, frequent 



