1963] 



BOTANY OF THE GUAYANA HIGHLANDS PART V 



139 



tiliter dimorphica; filamenta 4-5 mm longa apicem versus modice glanduloso- 

 setulosa; antherarum thecae 2-2.2 mm longae non vel vix attenuatae, connectivo 

 0.5-0.7 mm prolongate* basi in appendice cordiformi 0.5 X 0.7 mm expanso. Stylus 

 7.5 X 0.5 mm basim extremum versus sparse glanduloso-setulosus ; ovarium 4- 

 loculare densissime cum glandulis sessilibus obsitum et apicem extremum versus 

 glanduloso-setulosum. Capsula 4-valvata ; semina ca. 0.8 X 0.4 mm vix foveolata. 



Type. Petals pinkish white, filaments and connective bronze, anthers yellow, 

 frequent in lower part of lower northwest cumbre, Churi-tepui (Muru-tepui), 

 Chimanta Massif, Edo. Bolivar, Venezuela, alt. 2100-2200 m, 24 Jan 1953, John 

 J. Wurdack 34177 (holotype US 2342486). Paratypes. VENEZUELA, Edo. 

 Bolivar. Chimanta Massif: Churi-tepui, alt. 2250-2300 m, Wurdack 34208; 

 Central Section, alt. 1900 m, Steyermark & Wurdack 568; Abacapa-tepui, alt. 

 1800-2000 m, Steyermark 74855; Apacara-tepui, Car dona 1555; Acopan-tepui, 

 alt. 2100 m, Cardona 2302. Uaipan-tepui : alt. 1900 m, Car dona 2065. Auyan- 

 tepui : alt, 2100-2200 m, Tate 1148, Cardona 2652. 



M. aspera has rufous hairs on the lower leaf surfaces, young stems, and in- 

 florescence, as well as sepals 3-5 mm long and anthers 3-4 mm long. 



Macairea cardonae Wurdack, sp. now Fig. 64, a-d. 



M. lanatae Gleason affinis sed foliis 7-nervatis stylo glanduloso-setuloso. 



Frutex 0.5-3 m ; ramuli petiolis inflorescentiis hypanthiisque dense longo- 

 strigulosi vel brevistrigosi pilis gracilibus glanduliferis. Petioli 0.5-1.5 cm; 

 lamina 3.5-7 X 3-4.5 cm ovata apice late acuta vel obtusa basi vix (2-5 mm) 

 cordata- crasse coriacea, supra dense tuberculato-strigosa setis glanduliferis, 

 subtus in venis dense brevi-setulosa et in superficie densissime setulosa pilis 

 gracilibus plerumque glanduliferis, 7-nervata nervis primariis secundariisque 

 supra impressis et subtus leviter elevatis nervis tertiariis reticulatis. Panicula 

 3-5 cm longa lataque subpauciflora terminalis ; flores 3-5 mm pedicellati, 

 bracteolis 3-5 X 1.5-2 mm ellipticis ca. 2 mm infra hypanthium insertis. Hy- 

 panthium 3-3.5 mm longum ; calycis tubus 0.5 mm longus, lobis 3-3.5 mm longis 

 oblongis acutis basi remotis. Petala 14-14.5 X 13-14 mm suborbicularia apice cum 

 setulis glanduliferis paucis ciliolata. Stamina vix dimorphica ; filamenta 7 vel 

 5-5.5 mm longa modice glanduloso-setulosa ; antherarum thecae 3.7-4 vel 3.5-3.7 

 mm longae apice vix subulatae, connectivo sub loculis 0.3-0.5 vel 0.7 mm pro- 

 longate, apoendice cordato-rotundata 1 X 0.8-0.9 mm. Stylus 7 X 0.4 mm modice 

 glanduloso-setulosus; ovarium 4-loculare apice dense setuliferum setulis gland- 

 uliferis ad 1.5 mm longis. Capsula 4-valvata ; semina 0.7-0.8 X 0.5 mm minutis- 

 sime denseque muriculata. 



Type. Shrub 3 m, flowers pink, cumbre of Acopan-tepui, Edo. Bolivar, Ven- 

 ezuela, alt. 2200 m, Oct 1947, F. Cardona 2283 (holotype US 1932223). Paratypes. 

 VENEZUELA, Edo. Bolivar. Auyan-tepui: Cardona 256. Chimanta Massif: 

 Apacara-tepui, alt. 1800-2000 m, Cardona 1975 (VEN), Steyermark 75844; 

 Churi-tepui (Muru-tepui), alt. 2250-2300 m, Wurdack 34220; central section, 

 Rio Tirica, alt. 1900-1950 m, Steyermark & Wurdack 604, 936. 



M. lanata has 5-nerved leaf blades and a glabrous style. Another close relative, 

 M. neblinae Wurdack, has 5-nerved basally obtuse leaf blades and many -flowered 

 inflorescences, as well as a glabrous or only basally sparsely pubescent style. The 

 Cardona collections of M. cardonae had previously been identified as M. multi- 

 nervia Benth. ; that species, however, has well-developed inflorescences, smaller 

 leaf blades, glabrous style, and predominantly 3-celled ovary. 



