1963] 



BOTANY OF THE GUAYANA HIGHLANDS PART V 



171 



Miconia phaeophylla Triana. 



M.-fendleriana Cogn. DC. Mon. Phan. 7: 822. 1891. 



There are no floral differences between the Peruvian and Venezuelan collec- 

 tions, and Guayana material easily bridges the obscure gap in leaf base char- 

 acters. Of more import is the distinctness from other related species. Flower-mery 

 in the group is a difficult character to use; M. puberula Cogn. seems predom- 

 inantly 4-merous (as shown also in the Flora Brasiliensis illustration, despite 

 Cogniaux' 5-merous description), while M. brevipes Benth. seems to be 4-5 (-6, 

 fide Gleason's notes )-merous. The poorly known M. kappleri Naud. and M. fragilis 

 Naud. are both suggestive of this relationship ; they apparently differ from the 

 pubescent species in the glabrous stem internodes and leaves and (except for 

 M. puberula) the smaller flowers. The Guayana complex, including the M. 

 minutiflora (Bonpl.) DC. alliance, may be tentatively keyed as follows: 



1. Young internodes and lower leaf surfaces glabrous. M. minutiflora and allies. 



1. Young internodes and lower leaf surfaces densely stellate-puberulous. 



2. Large anther thecae 1 mm long or less. M. puberula. 



2. Large anther thecae 1.5-2.5 mm long. 



3. Young stem hairs all sessile-stellate. M. phaeophylla. 



3. Young stem hairs in part stipitate-stellate. 



4. Lower leaf surface hairs ca. 0.4 mm diam and stipitate-stellate. M. rubiginosa. 



4. Lower leaf surface hairs ca. 0.25 mm diam and mostly sessile-stellate. M. brevipes. 



Specimens examined : 

 M. puberula. 



PERU, San Martin : King 3362, Spruce s.n. (isosyntype NY). 

 M. phaeophylla. 



PERU, San -Martin: Spruce 4861 (isotype NY). COLOMBIA, Vaupes: Cerro 

 La Campana, Schultes 5569 (NY). Huila : Hacienda Miraflores, Little 7307 (US). 

 VENEZUELA, Trujillo: Pittier 13134 (NY, US). Carabobo : Pittier 8019 (US). 

 Aragua: Llewelyn Williams 10243 (US), 10467 (US). Bolivar: Kavanayen, 

 Maguire & Wurdack 33990; Cerro Perro, Cardona 756 (US) ; Uaiparu, Bernardi 

 6566 (NY) ; Cerro Cotorra, Steyermark 86868 (US) ; Rio Asa, Steyermark 86732 

 (US). Amazonas: Savanna Hills, Cerro Duida, Tate 724 (NY); Esmeralda, 

 Llewelyn Williams 15320 (US), Maguire & Wurdack 34630. BRITISH GUIANA. 

 Utschi Falls, Kamarang River, Tillett & Tillett 45801. 

 M. brevipes. 



VENEZUELA, Amazonas: Pto. Ayacucho, Llewelyn Williams 13805 (US); 

 Sanariapo, Llewelyn Williams 13066 (US) ; Santa Barbara, Cowan & Wurdack 

 31597; Esmeralda, Llewelyn Williams 15329 (US), Maguire & Wurdack 34668, 

 Steyermark 57763 (NY). Bolivar: Cerro Perro, Cardona 750 (US); Cerro 

 Bolivar, Wurdack 34461 (var. longifolia Cogn. ? ; with large leaves, suggestive 

 of M. phaeophylla). BRITISH GUIANA. "Roraima," Schomburgk 934 (prob- 

 able isotype, fide Gleason, NY) ; Isherton, Rupununi River, A. C. Smith 2475 

 (US) ; 35 miles south of Mackenzie, Cowan 39258. 



Miconia arirambae Huber. 



No sheet of the type number, Ducke 8089, was found by curators at the larger 

 European herbaria (G, K, L, M, P, W), nor in the two Rio de Janeiro collections 

 (R, RB). Among the miscellaneous sheets at the Museo Goeldi is a sheet of this 



