1963] 



BOTANY OF THE GUAYANA HIGHLANDS PART V 



175 



5-nerved cordulate leaf blades; flowering material is required to establish the 

 firm identity of the Yutaje shrub. 



Tococa guianensis Aubl. var. orinocensis Wurdack, var. nov. 



Rami modice vel densiuscule setosi, pilis eglandulosis fuscis 5-10 (-15) mm 

 longis gracilibus laxis. Hypanthium modice vel densiuscule setosum, pilis fuscis 

 2-3 mm longis gracilibus eglandulosis. 



Type. Shrub 0.7-1.5 m, petals pink, fruit reddish, frequent along moist edge 

 of Sabana Huachapana, Rio Orinoco 30 km above Santa Barbara, Terr. Ama- 

 zonas, Venezuela, alt. 125 m, 15 Sep 1957, Bassett Maguire, John J. Wurdack, 

 <& William M. Keith 41468 (holotype US 2342579). Paratypes. VENEZUELA, 

 Terr. Amazonas. Rio Orinoco: just west of Santa Barbara at junction of Rios 

 Orinoco and Ventuari, alt. 125 m, Cowan & Wurdack 32028; Sabana de Atures, 

 alt. 120 m, Llewelyn Williams 15935 ( VEN). Rio Ventuari: Sabana de Moyo on 

 south bank of Rio Ventuari 15 km above mouth, alt. 150 m, J. S. Level 33. Edo. 

 Bolivar. Rio Orinoco: morichal 2 km east of river near Cerro Gavilan (Cerro 

 Carichana), alt. 100 m, Wurdack & Monachino 39937. 



Nowhere in T. guianensis or its satellite species (T. spruceana Cogn., T. 

 occidentalis Naud., T. latifolia Naud., T. formicaria Mart.), including the more- 

 recently described taxa discussed by Gleason (Bull. Torrey Club 58: 250-251. 

 1931) does the above diagnostic combination along with very large spheroidal 

 formicaria occur; setulose hypanthia (the hairs usually gland-tipped) are found 

 in various units. The variability loci in the geographically discontinous rassen- 

 kreis are both too involved and incompletely studied for elaboration here (i.e. 

 both time and complexit} T have me stymied for now). 



Tococa egensis Naud. 



T. glandulosa Gleason, Bull. Torrey Club 58: 252. 1931. 



King 424 and Asplund 14596 cannot be distinguished from the Poeppig collec- 

 tions {2549, 2790, Addenda 180, W) ; all have (despite Cogniaux' description) 

 styles moderately puberulous with slender hairs and the minutely conical ovary 

 apex with about 10 delicate glandular setae 1-1.5 mm long. To T. egensis, I have 

 tentatively referred a large series of Amazonas (Venezuela) specimens with 

 sparsely glandular-setulose external calyx teeth up to 4 mm long and sparsely 

 glandular-setose hypanthia. I seriously doubt that T. longisepala Cogn. should 

 be maintained apart from T. egensis; although the Spruce type collections 

 (K, W) show no trace of hypanthial or calycine setae, there seem to be no other 

 differences from the Venezuelan material. The establishment of subspecific status 

 for T. longisepala under T. egensis would require critical examination of prior- 

 named species — T. coronata Benth. and T. truncata Benth. — as well as the species 

 of Tococa Sect. Anaphysca with puberulous styles and ovaries. 



Tococa obovata Gleason subsp. neblinensis Wurdack, subsp. nov. 

 Folia supra bullis setif eris pustulata subtus foveolata. 



Type. Shrub to 1 m, stems and lower leaf surface red-tinged, hypanthium 

 maroon, petals outside 1/2-white and 1/2-pink, inside pink, frequent at Cumbre 

 Camp, Cerro de la Neblina, Terr. Amazonas, Venezuela, alt. 1800 m, 12 Nov 1957, 

 Bassett Maguire, John J. Wurdack, & Celia K. Maguire 42055 (holotype US 

 2342602). Paratypes. VENEZUELA, Terr. Amazonas. Cerro de la Neblina: 



