1963] 



BOTANY OF THE GUAYANA HIGHLANDS PART V 



185 



C. flexuosa has reniform or cordate small leaves, sinuate-margined large leaves, 

 stella te-puberulent and setose branches, and sessile (with pedicel less than 0.2 mm 

 long) flowers. Of the 4-merous species of Sect. Calophysoides, only C. semijuga 

 (Gleas.) Wurdack shows as complete a degree of leaf abortion as C. alternifolia; 

 that species, however, has eciliate leaf blades, well-developed inflorescences, 

 glabrous hypanthia, stamen connectives with a dorsal appendage, and costate 

 fruit, 



Clidemia pakaraimae Wurdack, sp. nov. 



Sect. Sagraea. C. eymiferae Donn. Sm. affinis sed foliis maioribus paniculae 

 ramis crassis hypanthiis non stellulato-furfuraceis ovario omnino infero. 



Rami primum quadrangulares demum compresso-quadrangulares infra nodos 

 tumidi vesicis binis vel solitariis 1-2 cm longis 0.5-1 cm altis hypanthiis petioliis- 

 que non setosis vel primum sparsissime glanduloso-setosis demum glabratis 

 petiolis foliis juvenilibus inflorescentiis hypanthiisque dense appresseque resinoso- 

 furfuracei tarde laeves. Petioli 6-12 cm; lamina 15-28 X 9-21 cm ovata apice per 

 1.5-2.5 cm graditer acuminata et apiculata basi 0.5-2.5 cm cordata tenuiter 

 coriacea, supra et subtus non setulosa ad margines integra et ciliolata pilis 

 appressis ca. 0.5 mm longis 0.3-0.5 mm inter se distantibus, 7- vel sub-9-nervata 

 ncrvis secundariis 5-8 mm inter se distantibus nervulis supra invisis subtus 

 densiuscule reticulatis. Panicula 4—5 cm longa lataque pauciflora ramulis robustis 

 compressis 1-2 mm crassis; bracteae bracteolaeque 1-3 mm longae triangulares 

 acutae persistentes ; pedicelli 1-3 mm longi robusti ; flores 4-meri. Hypanthium 

 4 mm Ion gum ; calycis tubus 1 mm altus, lobis interioribus 0.5 mm altis oblatis 

 apice rotunda! is, dentibus exterioribus 0.8 mm eminentibus linearibus crassis 

 hebeti-ai-utis vel setuloso-apiculatis ; torus non setulosus. Petala 7 X 3.3-3.5 mm 

 obovato-oblonga apice truncato-rotundata. Stamina isomorphica glabra ; filamenta 

 4.5-4.8 mm; antherarum thecae lineares 2.1 mm longae apice retusae poro 0.2 

 mm diam dorsaliter inclinato, connective basi 1-1.2 mm prolongato simpliciter 

 articulato. Stylus 6.5 X 0.5-0.7 mm glaber ; stigma 0.8 mm diam capitulatum ; 

 ovarium 4-loculare omnino inferum glabrum. 



Type. Shrub to 3 m, flowers creamy white, locally frequent in wet forest, 

 Merume Mountain, Partang River, upper Mazaruni River basin, alt. 1140 m, 

 1 Jul 1960, Stephi n S, Till* ft, Carolyn L. Tillett, d- Rufas Boyan 43949 (holotype 

 US 2343841 . 



C. cymifera has leaf blades only 5-12 cm long, slender cyme branches only 

 0.5 mm diam at most, and a 1/3 superior ovary; the stems, lower leaf surface 

 veins, inflorescence, and hypanthium are stellulate-furfuraceous. Otherwise, the 

 Guatemalan species is quite similar in qualitative floral details. C. ampla Cogn. 

 (which is very similar to, if not synonymous with C. grandifolia Cogn.) is more 

 distantly related ; the young stems and petioles are densely lax-strigulose with 

 roughened hairs, the bracts are setulose-ciliolate, the flowers are much smaller, 

 and the well-developed ovarian collar is apically setulose. I do not know if the 

 nodal swellings of C. pakaraimae are genetic or pathologic — the occasional uni- 

 lateral development suggests the latter. Certainly these inflations differ in appear- 

 ance from the cauline formicaria of C. tococoidea (DC.) Gleason. 



C. pakaraimae does not suggest any affinity with C. coriacea (Naud.) Cogn. 

 and its close relative, C. duidae Gleason ; both of these cumbre species have 

 terminal inflorescences and probably should be transferred to Miconia, perhaps 

 among the 4-merous species of Sect. Octomeris. I have continued to use the later 



