226 



MEMOIRS OF THE NEW YORK BOTANICAL (JARDEX 



[VOL. 10 



of stipule, no correlation can be found in these characters between the plants of 

 Roraima (where Chalepophyllum speciosum was described) and those of Auyan- 

 tepui (where Chalepophyllum coriaceum was described), or in those plants col- 

 lected on Ptari-tepui, Chimanta Massif, and Serra do Sol. Gleason described his 

 C. coriaceum as having the acumen of the stipules "erect, appressed" and "2-4 

 long" with internodes "mostly 1.5-3 cm long" (loc. cit. p. 192), contrasting this 

 with C. speciosum which, according to him, had stipules ' ' much longer and more 

 or less spreading" and internodes "only 1-10 mm long." Unfortunately, these 

 differences, likewise, do not hold, variations in internode length and stipule length 

 occurring on the same plants. Where the specimen has been collected from elon- 

 gated vegetative shoots or from a portion of the main thicker branch, the inter- 

 nodes show, for the most part, greater lengths than those of the short lateral and 

 upper terminal shoots. There is as much crowding of the internodes on flowering 

 and fruiting branches of plants from Roraima (type locality of Chalepophyllum 

 speciosum) as from those of Auyan-tepui (type locality of C. coriaceum). In the 

 original plate illustrating C. speciosum the internodes are shown varying from 

 short and crowded (4-5 mm long) on the branch at. the lower side of the repro- 

 duction to longer (8-11 mm long) on the branch at the right side of the same 

 illustration. All degrees of variation are exhibited on collections from Chimanta 

 Massif, the collections of Steyermark & Wurdack 1004 having prolongations of 

 the stipules 2-2.5 mm long, while Steyermark 75781 has stipule prolongations 

 of 5-6 mm long on elongated vegetative shoots. Topotype material of C. speciosum 

 from Roraima (represented by Tate 388, Steyermark 58860, and Mc Connell & 

 Quelch 653) shows internodes from 3-8 mm long and stipule tips from 3-5 mm 

 long, whereas holotype and topotype material of C. coriaceum from Auyan-tepui 

 (represented by Tate 1179, Car dona 2669, Foldats 2603, and Vareschi & Foldats 

 4858) has internodes ranging from 3-20 mm long and stipule tips from 2-4 mm 

 long. Collections from Chimanta Massif have internodes ranging from 3-14 mm 

 long and stipule tips from 2-6 mm. Collections from Ptari-tepui have internodes 

 3-5 mm long (Steyermark 59740) to 10-15 mm long (Steyermark 59801) and 

 stipule tips from 2-3.5 mm long. 



As regards length of corolla-tube and length and shape of corolla-lobes, great 

 variation is encountered. On topotj^pical material from Mount Roraima of 

 Chalepophyllum speciosum the corolla-tube varies from 45-90 mm long, the 

 corolla-lobes from 25-50 mm long and mainly 7-12 mm broad (4 mm in dried 

 wrinkled material), while in topotypical material from Auyan-tepui of C. 

 coriacefUm the corolla-tube varies from 70-100 mm long, the corolla-lobes from 

 20-36 mm long and 7-12 mm broad. Specimens from Chimanta Massif range from 

 corolla-tube lengths of 65-115 mm long and corolla-lobes 25-48 mm long and 

 7-15 mm broad. From Ptari-tepui the length of the corolla-tube measures 85-90 

 mm long, the corolla-lobes 35 mm long by 7-8 mm broad. Specimens from Serra 

 do Sol (Maguire & Maguire 40392) show the greatest extremes in width of 

 corolla-lobes, these having a width of 14-19 mm. The length of 40-47 mm in this 

 latter collection is matched by a Wurdack 34209 collection from Chimanta Massif 

 having corolla-lobes 42-48 mm long but a width of 9-11 mm. There is too much 

 intergradation in corolla-lobe width to segregate the Serra do Sol collection. So 

 far as corolla-lobe w 7 idth, there are overlapping and non-correlating variations 

 in width among the range of material passing as Chalepophyllum speciosum and 

 C. coriaceum. Also, in many specimens the corolla-lobes are not pressed equally 

 flat, resulting in some shrinkage with measurements of width less than in well- 

 pressed or living material. 



