1963] 



TAXONOMY OF LUZULA SUBGENUS PTERODES 



289 



The blades of the culm leaves are over 4 cm long and from 1-2 (rarely 3) mm 

 wide. This is a wide ranging variety that extends from the Canary Islands, 

 east through northwest Africa to Greece and north into Germany and southern 

 England (Fig. 2). 



Type. Mr. Edw. Forster, jun. In a wood opposite the south front of Hasod 

 house, Cardiganshire, (not seen.) 



Kepresentative Specimens: ALBANIA: Alston $ Sandwith 1744 (K). AUSTEIA: Sauter 

 1049 (GH, US). BELGIUM: HeurcJc $ Martinis 290 (K) ; A. Lelievre 180 (GH, LISU). BUL- 

 GAEIA: C. K. Schneider 537 (K). CANARY ISLANDS: E. Asplund 166 (K) ; J. Ball 

 (March, 1888) (GH) ; E. Bourgeau 1554 (K) ; B. P. Murray (May 7, 1902) (K) ; Perrandiere 

 (April 4, 1855) (GH, US). CORSICA: M. F. Spencer (May 30, 1906) (GH). ENGLAND: E. 

 Armitage 1656 (K) ; A. Bennett 1002 (US) ; C. B. Billups 1549 (K) ; C. B. Clarke (May 28, 

 1888) (K), (July 9, 1900) (K), (June 17, 1905) (K) ; J. S. Gamble 18768 (K) ; Jackson $ 

 Green 1656 (K) ; J. E. Little 2 (K) j N. Y. Sandwith (June 5, 1921) (K), (June 18, 1927) 

 (K) ; V. S. Summerhayes 1936 (K) ; W. B. Waterfall 1352 (K) ; H. C. Watson 1171 (US) ; 

 D. P. Young 3992 (K). FRANCE: E. Chabert 2639 (LISU); W. P. Conant (Oct. 10, 1879) 

 (GH); A. Ditiglize (May 30, 1867) (US); H. H. litis 3048 (US); S. E. Lassimonne (April, 

 1892) (US); F. Baine (March 21, 1907) (GH) ; A. Songeon (May 12, 1857) (GH) ; Vahl 

 (April, 1825) (US). GERMANY: Kneucker 55 (GH, K, US) ; F. Schultz 154 (K, LISU); F. 

 von Speissen (May 10, 1879) (US), (March 24, 1880) (US). HUNGARY: L. Bichter 55 (GH, 

 US). ITALY: J. Ball (April, 1842) (GH), (March 23, 1862) (GH), (June, 1863) (GH), 

 (May 9, 1866) (GH), (May 6, 1867) (GH) ; A. Beguinot 411 (GH, K) ; E. F. Paoletti (March, 

 1900) (GH); H. Boss 687 (GH). PORTUGAL: J. Daveau (March, 1882) (LISU); M. Fer- 

 reira 536 (LISU); A. de C. Freire 183 (LISU); A. Moller 56 (LISU, US), (March, 1879) 

 (LISU); C. Bomariz 39 (LISU), 860 (LISU), 869 (LISU); M. Winkler (May 8, 1876) (K). 

 SPAIN: N. Y. Sandwith 5258 (K), 5286 (K) ; F. Spennen (March, 1920) (K) ; Willkomm 587 

 (GH). SARDINIA: P. Thomas (1828) (US). SWITZERLAND: Favrat # Barley (May, 

 1872)' (K); B. Masson (May, 1888) (US). YUGOSLAVIA: C. Baenitz (April 21, 1898) 

 (US) ; Jackson # Turrill 72 (K). 



Hybrids. The only naturally occurring hybrid that has been found during 

 this study is Luzula forsteri X L. pilosa. This hybrid was first described as a 

 species (Luzula borreri Bromf. ex Bab.) by Babington (1851), and later treated 

 as a variety of L. pilosa by Syme (1872). Buchenau (1890) realized that this 

 taxon was a hybrid and later botanists have treated it in this same manner. 



2b. Luzula forsteri (Smith) DC. in Lam. & DC. var. rhizomata Ebinger, var. 

 nov. 



A va^. forsteri rhizomatibus 2-4 cm longis ; caulibus 20 cm altis ; laminis 

 foliorum caulinorum 2-4 mm longis differt. 



This variety is similar to variety forsteri, but differs in that rhizomes are 

 present. These are to 4 cm long and 2 mm in diameter. The plant is 20 to rarely 

 30 cm tall, and the culm leaves do not exceed 4 cm in length and 1-2 cm in 

 width. This variety is restricted to the eastern Mediterranean Region (Fig. 2). 



Type. Davis 21536: Turkey, Cankiri Providence, Ilgav District, Yaylacik; 

 1000 meters in Pinus nigra forest; June 5, 1954. (K.) 



Kepresentative Specimens: IRAN: H. F. Mooney 6625 (K). LEBANON: O. Polvnin 5325 

 (K). LESBOS: C. N. Goulimy (April 12, 1957) (K). RUSSIA: (Crimea): C. Golde 1092 

 (K). TURKEY: Davis 21536 (K— HOLOTYPE) ; D. Eeldreich 3550 (K). 



Hybrids. None have been reported and none were found during this study. 



Luzula forsteri is a wide ranging species that is very distinct and easily dis- 

 tinguished from any other member of the subgenus. Morphologically it is more 

 closely related to L. plumosa, from which it differs in the lack of a large hooked 

 caruncle on the seed, and the presence of a mucro on the callose tip of the leaf. 



