1963] 



TAXONOMY OF LUZULA SUBGENUS PTERODES 



299 



mosa by its spreading to reflexed pedicels. This variety is generally restricted 

 to the islands of Japan, but is occasionally found in Korea (Fig". 7). 



Type. Savatier 3368: collected in the mountainous province of Senano (now 

 called Nagano). Saba, Japan, (not seen.) 



Kepresentative Specimens: JAPAX: Hokkaido: P. Faurie 238 (K), 3764 (K) ; Maximo- 

 wicz (May, 1861) (K). Honshu: Beattie 4' Eurihara 10738 (GH, US) ; I. Bate (May 18. 

 1958) (TXS); P. Faurie 252 (K), 1014 (K) ; M. Furuse (April 25, 1954) (S), (June 5. 

 1955) (S, TXS); G. Hashimoto (Aug. 7, 1931) (TXS); E. Iishiba (May 9, 1926) (S) ; C. 

 Eozada 9 (TI) ; L Matsumoto (May 5, 1955) (TXS); M. Mizushima (June 16, 1951) (S), 

 (June 27, 1952) (S, TI) ; G. Murata (May 3, 1956) (S) ; Y. Satake 190 (TI) ; Satake $ 

 Okuyama (May 23, 1942) (TXS), (May 24,*1942) (TXS) ; E. Shiota 3378 (GH), 5533 (GH), 

 5685 (GH), 9629 (GH), 17203 (GH) ; I. Sono (June 1, 1907) (TXS); M. Togasi 1035 (K, 

 S, US) ; Togasi ^ Eoidzumi 730 (K, S, US) ; T. Tsuchihashi (June 3, 1926) (TXS). Kyushu: 

 E. Ichikawa 218 (US); E. Yamashita (June 12, 1931) (TI). KOREA: C. T. Hyoun 1583 

 (SKK) ; Taquet 2143 (K). 



Hybrids. Xo naturally occurring hybrids have been reported, and none were 

 found during this study. The only taxon that it is possible for variety brevipes 

 to hybridize with is Luzula rostrata. Hybrids were not found, however, which 

 is probably the result of the difference in flowering time of the two taxa. It is 

 also possible that an ecological barrier exists, but this could not be determined 

 with certainty. From the data that was present on the herbarium sheets it was 

 determined that variety brevipes blooms during April and May, while L. rostrata 

 blooms in June and July. Also, L. rostrata is restricted to mountainous regions 

 while variety brevipes grows at low altitudes. There were, however, a few speci- 

 mens of this variety that bloomed later in the season while other specimens were 

 found growing at fairly high altitudes. Thus the opportunity for hybridization 

 may sometimes occur. 



Morphologically this species is intermediate between the European species 

 and the Asiatic species Luzula rostrata. L. plumosa var. phi mosa is similar to 

 L. forsteri in that the pedicels are erect, the flowers are essentially the same size, 

 and the capsules equal to slightly exceed the perianth. It can be separated 

 readily from this European species, however, since the leaves are wider, there 

 is no mucronate projection extending from the callose tip of the leaf, and the 

 caruncle is much larger. In contrast, var. reflexa is morphologically very close 

 to L. pilosa. It can be separated from this species by the wider and longer culm 

 leaves, the slightly longer perianth and the anther length which equals to slightly 

 exceeds the filaments. Variety brevipes, in contrast, is more similar to L. rostrata 

 than the European species. However, it can be distinguished from L. rostrata by 

 its narrower culm leaves that do not taper abruptly to the callose tip, the longer 

 perianth segments and the strongly compound inflorescence with spreading to 

 reflexed pedicels. 



The nomenclature of Luzula plumosa var. plumosa is very simple, and only 

 three synonyms are listed. Two of these represent transfers of the specific epi- 

 thet plumosa, either to varietal status or to another generic name that is now 

 considered a synonym of the genus Luzula. The third is a herbarium name, J un- 

 cus plumosus, which was listed as a synonym of L. plumosa. Variety brevipes, 

 in contrast, has numerous synonyms which are a result of the large amount of 

 variation found in this variety. It has been generally considered to be allied to 

 L. plumosa, and in the most recent flora of Japan is referred to as L. plumosa 

 var. macrocarpa (Buchenau) Ohwi (1953). This variety name was taken from 

 L. rufescens var. macrocarpa Buchenau. The type of this name was examined 



