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MEMOIRS OF THE NEW YORK BOTANICAL GARDEN 



[vol. 11 



pachyglossaceae Fulford, fam. nov. 

 Lophocoleaceae Herzog & Grolle, Revue Bryol. Lichenol. 27:162. 1959. 



Caules foliosi erecti, radiales, ramis basilaribus stolonoideis, fusci; ramis 

 foliosis flagelliformibusve in axillis foliorum amphigastriorumque. Folia amphi- 

 gastriaque transversa, ex ovatis ad breviter rigida, pluristratosa omnino (in 

 specie uniea ad medium). Inflovescentia masculina terminalis turn intercalaris ; 

 inflorescentia feminea terminalis, innovationibus subfloralibus praesentibus vel 

 absentibus; bracteis braeteolisque foliis similibus, pluristratosis. Perianthium 

 supra 3-5-earinatum. 



Leafy stems erect, with stolon-like branches at the base, radially symmetric ; 

 branches leafy, more rarely flagelliform, axillary from the leaves and under- 

 leaves. Rhizoids from the scale leaves of the flagelliform branches. Line of leaf 

 insertion transverse, the ends slightly decurved. Leaves and underleaves scarcely 

 different, pluristratose throughout or in the lower half. Male inflorescence 

 terminal becoming intercalary on the stem or leafy branch. Female inflorescence 

 terminal on the stem or branch, with or without subfloral innovations, the bracts 

 and bracteoles like the leaves and underleaves, larger, pluristratose. Perianth 

 three-lobed or divided above, 3-5-keeled, unistratose. Sporophyte not seen. 



Type genus : Pachyglossa Herzog & Grolle. 



The family is monogeneric ; the distribution is Antarctic. 



Pachyglossa, Herzog & Grolle, Revue Bryol. Lichenol. 27: 150. 1958. 



Pachyglossa Herzog, Rev. Bryol. Lichenol. 21: 256. 1952. (nomen nudum.) 



Plants of small to medium size, erect, with stolon-like branches at the base, 

 radially symmetric, greenish-brown to dark brown, in tufts or among other 

 bryophytes. Rhizoids from the scale-like leaves of the flagelliform branches and 

 the rhizome. Line of leaf insertion transverse, the ends slightly decurved. 

 Leaves and underleaves ligulate to ovate, the apex blunt, refuse or very shortly 

 bifid, pluristratose throughout (in one species only to the middle). Plants 

 dioicous. Male inflorescence terminal, becoming intercalary on the stem or leafy 

 branch, the bracts and bracteoles in few series, similar, like the leaves and 

 underleaves, larger, in some with a large marginal tooth; antheridia in the 

 axils of the bracts, the stalk uniseriate. Female inflorescence terminal on the 

 stem or a leafy branch, without or with one or more innovations, the bracts and 

 bracteoles in two or three series, similar to the leaves and underleaves, larger, of 

 two or three layers of cells; archegonia less than 20. Perianth 3-keeled, deeply 

 3-parted above, or 4- or 5-plicate with the mouth 3-lobed. Sporophyte not seen. 



Type species, J ungermannia ienacifolia Hooker f. & T. Taylor. 



KEY TO THE SPECIES 



1. Leaves of two to several layers throughout; superficial leaf-cells 15-20 X 13-35 n. 

 2. Leaves ligulate to ovate, obtuse, entire, refuse or very shortly bifid. 



3. Leaf cells uniformly thick-walled; stems in transverse section with several layers 



of very thick-walled cortical cells, (not seen in S. America) 1. P. tenacifolia. 



3. Leaf -cells with thin walls and small trigones; stem in transverse section with 



only one layer of thickened cortical cells. 2. P. dissitifolia. 



2. Leaves broadly ovate, obtuse and shortly bifid; perianth 4-5-plicate, the mouth 



irregularly dentate and short-ciliate. 3. P. fissa. 



1. Leaves of two layers of cells only in the lower part, the upper part unistratose; 



leaf -cells 10-16 X 10-20^; leaves obovate, undivided. 4. P. spegazziniana. 



