1962J 



LEAF'S IIKPYT1CAE OF LATIN AMERICA — I'ART I 



85 



Herzog, T. & Grolle, R. 1958. Was ist Pachyglossal Revue Bryol. Lichenol. 27: 147-165. 

 f. 1-5. 



Hattori, S. & Mizutani, M. 1958. Pachyglossa iristicha Herz. and its relationships. Jour. 

 Jap. Bot'. 33: 359-363. /. 1, S. 



herbertaceae K. Midler in Rabenhorst, Krypt.-Fl. 6: 557. 1954. emend. Fulford 

 & Hatcher, Bryologist 61 : 284. 1958. 

 Ptilidiaceae auctt. p.p. 



Leafy stems erect or pendent, tending to be radially symmetric, brown, 

 irregularly branched (in Herbert a the stems arising from a creeping caudex 

 with scale-like leaves) ; branches both terminal (Fntllania type) and intercalary 

 in orgin, the intercalary branches ventral, axillary, or on the stem at the ventral 

 side of the leaf. Line of leaf insertion transverse, or oblique with the leaves 

 incubous. Leaves bifid or trifid, with lanceolate divisions. Underleaves like the 

 leaves, rarely much smaller. Male inflorescence terminal but becoming inter- 

 calary on the stem or branch, radially symmetric; antheridia in the axils of 

 both the bracts and the bracteoles, the stalk of two rows of cells. Female inflor- 

 escence terminal on the stem or a leafy branch, without or with one or two 

 innovations, the bracts and bracteoles essentially like the leaves and under- 

 leaves. Perianth long, contracted above, three- to six-plicate, divided to one- 

 fourth or more into six to nine entire or toothed segments. 



Type genus : JI< rb< via S. F. Gray. 



Key to the Genera 



Leaves and underleaves with a well-marked vitta of elongate cells; leaf cells thick- 

 walled, with large, knot-like trigones and intermediate thickenings. 2. Herberta. 



Leaves and underleaves without a vitta; leaf -cells quadrate to hexagonal in outline, 



thin-walled, without trigones. 1. Triandrophyllum. 



Triandrophyllum Fulford & Hatcher, Bryologist 64: 349. 1961 [1962]. 

 Isotachis auctt. p.p. 



Plants large or small, ascending to erect, simple or irregularly branched. 

 Lateral branches of the Fntllania type with the half-leaf at the dorsal base, 

 leafy, ventral branches frequent, intercarlary, from the axils of the underleaves, 

 leafy or flagelliform ; stem in transverse section 12-16 cells in diameter, the 

 cells of the cortex in one or two layers, small, thick-walled, those of the medulla 

 large with thinner walls. Line of leaf insertion oblique, recurved at the dorsal 

 end, the leaves incubous. Rhizoids not seen. Leaves asymmetric, bifid or trifid to 

 one-fifth or one-half of their length ; segments acute, triangular, the margins 

 dentate or spinose or entire ; leaf-cells small, round-quadrate to hexagonal, thin- 

 walled, the trigones minute or absent, the cuticle smooth to verruculose-papil- 

 lose. Underleaves as large as the leaves, similar, or smaller with the margins 

 abundantly toothed. Plants dioicous. Male inflorescence terminal becoming 

 intercalary on the stem or the branch, the bracts and bracteoles in eight or more 

 series, similar to the leaves and underleaves, larger, concave ; antheridia large, 

 globose, two or three in the axils of both the bracts and the bracteoles, the stalk 

 of two or more rows of cells. Female inflorescence terminal on the stem or a 

 branch, the bracts and bracteoles in two or three series, similar to the leaves 

 and underleaves, slightly larger and with more marginal teeth. Perianth ob- 



