180 



MEMOIRS OF THE NEW YORK BOTANICAL GARDEN 



[VOL. 11 



Plants of small to medium size, green, pigmented with brown, in tufts or 

 mats or among other bryophytes; stems ascending to erect, at least 2 cm long, 

 irregularly branched, the lateral branches leafy, diverging at a wide angle, the 

 ventral branches leafy or flagelliform, of the Acromastigum and ventral inter- 

 calary types; stem in transverse section with a unistratose cortical layer of 

 12 large cells similar to those of the medulla. Line of leaf insertion transverse 

 to slightly oblique, the leaves succubous. Leaves distant, spreading, cuneate, the 

 lateral margins nearly straight, entire, the base nine rows of cells wide, bifid or 

 trifid to about half of their length; segments equal or unequal, divergent, 

 triangular from a 4-6-celled base, ending in an acute tip of one or two cells; 

 leaf-cells 36-60 X 15-23 p., the walls thickened, the cuticle verruculose. Under- 

 leaves large, about half as large as the leaves, mostly cuneate, trifid (rarely 

 bifid) to one-third of their length, the segments divergent. Branch leaves and 

 underleaves a little smaller, more often only bifid. Male inflorescence on a short 

 ventral sexual branch, the bracts and bracteoles in 25 or more series, imbricate, 

 the bracts concave, bifid, often with a tooth on the lateral margins, the bracteoles 

 smaller, plane. Female inflorescence and sporophyte not seen. PI. 39. Fig. 1, a-f. 



Habitat: Not known. 



COLOMBIA: Bogota, "In desertis Paramo de San Fortunato," 2900 m, Lindig (type G). 

 The species is also reported from Costa Rica (Stephani. 1893) and Colombia (Herzog, 

 1934a). 



Lepidozia (Dumortier) Dumortier, Recueil Obs. Jungerm. 9. 1835. 4 

 Pleuroschisma sect. Lepidozia Dumortier, Syll. Jungerm. 69. 1831. 



Mastigophora C. G. Nees, Nat. Eur. Leberm. 1: 95, 101. 1833. Non Nees, Nat. Eur. 



Leberm. 3 : 85. 1838. 

 Herpetium sect. Lepidozia C. G. Nees, Nat. Eur. Leberm. 3 : 85. 1838. 

 Lepidozia subg. Eulepidozia Spruce, Jour. Bot. London 14: 164. 1876. 

 Lepidozia subg. Ptilolepidozia Spruce, Jour. Bot. London 14: 164. 1876. 



Plants in whitish, yellow-green, greenish-brown, or brown mats or tufts, often 

 pendulous or among other bryophytes; stems filiform to robust, 0.5-10 cm long, 

 pinnate or bi-tripinnate, the lateral branches usually of limited growth, of the 

 Frullania type with the dorsal half-leaf ovate, bifid, often becoming flagelliform 

 in the outer part, the ventral branches axillary, intercalary, flagelliform, occa- 

 sionally leafy, or very short, sexual. Stem in transverse section rounded, usually 

 pigmented with brown, little differentiated. Rhizoids colorless, the tips enlarged or 

 variously branched, from the scales of flagelliform branch tips, more rarely from 

 the bases of the underleaves. Line of leaf insertion straight to slightly curved, 

 subtransverse, oblique or almost longitudinal. Leaves incubous, alternate, sym- 

 metric or asymmetric, quadrate, rectangular, broadly cuneate or ovate-truncate, 

 the dorsal margin convex from a straight to cordate base, the margins of the 

 lamina entire or toothed, incised or ciliate, quadrifid (in some species with five 

 or six segments), the segments triangular; leaf-cells quadrate-hexagonal, the 

 walls uniformly thickened, the trigones when present minute. Underleaves trans- 

 verse, nearly as large as the leaves to much smaller, the segments triangular to 

 uniseriate, the margins entire, incised, toothed, or ciliate. Branch leaves and 

 underleaves larger or smaller. Plants dioicous, a few species monoicous, the 

 sexual branches short, ventral, axillary-intercalary. Male branches catkin-like 

 (rarely becoming intercalary on a leafy stem), the bracteoles plane, the bracts 

 concave, 2-5-parted above, diandrous (?); antheridia large, spherical, the stalk 



4 This revision has followed in a large part the work of Dr. J. Taylor, 1960. 



