1952] 



PLANTS COLLECTED IN ECUADOR 



77 



small, with much shorter calyx-lobes, corollas, and stamens. Another species of 

 this alliance, T. cardiopbylla Sleumer, also of Peru, has the leaves smaller than 

 those of the new species and the pedicels and comparatively large calyx densely 

 pubescent. 



Thibaudia martiniana A. C. Smith, sp. nov. 



Frutex ubique filamentis exceptis glaber, ramis elongatis, ramulis robustis 

 fuscis obtuse angulatis; petiolis crassis (3-5 mm. diametro) 13-20 mm. longis 

 manifeste angulatis; laminis coriaceis in sicco fusco-olivaceis ovato-ellipticis, 

 20-30 cm. longis, 10-17 cm. latis, basi obtusis et in petiolum decurrentibus, 

 apice ut videtur acutis vel breviter cuspidatis, margine integris et leviter re- 

 curvatis, pinnatinerviis, costa supra paullo subtus valde prominente, nervis 

 secundariis utrinsecus plerumque 3 curvatis supra conspicue impressis subtus 

 prominentibus, rete venularum supra subplano vel prominulo vel leviter impresso 

 subtus conspicuo; inflorescentiis axillaribus breviter racemosis circiter 10-floris 

 sed floribus saepe caducis, rhachi crassa superne angulata circiter 1.5 cm. longa, 

 bracteis caducis; pedicellis rugulosis sub anthesi 20-23 mm. longis superne valde 

 incrassatis basim versus bibracteolatis, bracteolis ovato-deltoideis obtusis cir- 

 citer 2 mm. longis, articulatione conspicuo; calyce coriaceo ruguloso cupuliformi- 

 cylindrico sub anthesi haud apophysato 8-10 mm. longo apice 6-10 mm. diametro, 

 tubo circiter 4 mm. longo, limbo suberecto quam tubo longiore inconspicue 5-dentato, 

 dentibus minute apiculatis haud 0.5 mm. longis, sinibus complanatis; disco carnoso 

 pulvinato; corolla crasso-carnosa cylindrica sub anthesi circiter 25 mm. longa et 

 basim versus 7 mm. diametro, superne gradatim angustata, lobis 5 deltoideis 

 obtusis circiter 1.5 x 2.5 mm.; staminibus 10 corollam subaequantibus, filamentis 

 liberis fuscis ligulatis circiter 5 mm. longis intus superne puberulis, antheris 

 circiter 23 mm. longis crassis basi obtusis, thecis in tubulos 7-8 mm. longos in- 

 ferne lateraliter connatos rimis elongatis dehiscentes gradatim transeuntibus; 

 stylo crasso sub anthesi leviter exserto, stigmate obscure papillose 



Pichincha: Along the road from Quito to Sto. Domingo de los Colorados (western 

 slope of the cordillera), 8,500-9,500 ft. elev., Jan. 15, 1945, Camp E-1717 (TYPE 

 US 1,989,049; dupl. NY) (on banks in soil; branches arching to 4 m.; leaves dark 

 green above, pale beneath, dull on both surfaces; pedicels red, the bracteoles 

 white, the "joint" green; calyx red; corolla pure white). 



At the suggestion of the collector, this new species is named for Or. William 

 E. Martin, of the University of California, in whose company the type specimen 

 was obtained. It is closely related only to T. pachypoda A. C. Smith, known from 

 low elevations near the coast of El Valle, Colombia, but it differs in its more 

 robust foliage, the leaf-blades being much larger and with more prominent second- 

 ary nerves. The new species also has shorter pedicels, a slightly shorter corolla, 

 and anthers with short tubules which are laterally adnate nearly to the apex. 



Another collection which may be mentioned here is Camp E-626 (NY only), 

 from El Oro, in Moro-Moro region about 21 miles west of Porto velo, 3,400-4,200 ft. 

 elev. (coarse shrub with branches to 3 m. long, in dense rain-forest). As compared 

 with the type of T. martiniana, no. 626, which is past anthesis, has shorter petioles, 

 pedicels up to 33 mm. long, and a shorter calyx-limb with more obvious lobes. 

 Without corroborating evidence from the corolla and stamens, I hesitate to expand 

 the concept of the new species to include this specimen, although it does not 

 suggest any other known species. 



Thibaudia parvifolia (Benth.) Hoer. Bot. Jahrb. 42: 275. 1909. 



Azuay: "Oriente" Border, crest of Eastern Cordillera, between Ona and the 

 Rio Yacuambi, 10,000-11,200 ft. elev., F. Prieto P-305 (shrub 2 m c ; leaves deep 



