1953] 



REVISION OF MACROLOBIUM 



329 



face, the wings glabrous beneath, the axis more or less pilose. Leaflets 13-44 

 mm. long, 8-15 nam. wide, the basal pair oval, the others oblong; puberulous on 

 the costa above, puberulous on the costa and sometimes also on the basal por- 

 tion of the blade beneath. Inflorescences 3.5"5 cm. long, axillary or terminal, the 

 peduncle 1-2 mm. long; pedicels 3-3.5 mm. long, puberulous; bracteoles 5-5.5 

 mm. long, 2.5 mm. wide, oblanceolate, externally puberulous basally and on the 

 costa, glabrous within. Hypanthium 5 mm. long on a 1 mm. stipe, sparingly pu- 

 berulous. Sepals 6-7 mm. long, 2-3 mm. wide, oblong, glabrous. Filaments about 

 12 mm. long, basally pilose. Ovary oblong-elliptic, pilose marginally, the lateral 

 surfaces glabrous, 4-ovulate; free portion of the gynophore 2.5 mm. long, pilose. 

 Fruit (immature) oblong, wider toward the apex, the margins sparsely pilosulose, 

 the carpophores about 10 mm. long; seeds 4. 



Type Collection: E, Delgado 299, "Camino antiquo de Caracas a La Guaira, 

 D. F.," Venezuela, July-Nov. 1939-40 (HOLOTYPE VEN, isotypes F, US). 



Additional Specimens: COLOMBIA: Cordillera Oriental, Dept. Norte de Santander, Sa- 

 rare region, confluence of Rios Cubugon & Cobaria, Not. 1941, Cuatrecasas 13204 (COL, 

 US). 



40e. Macrolobium colombianum var. colombianum. Figure 12. 



Outea "(?)" colombiana Britton & Killip, Ann. N. Y. Acad. 35: 166. 1936. 



Branchlets glabrous or the very young ones sparsely pilosulose. Stipules 5 

 mm. long, 1.5 mm. wide, subulate-lanceolate, acuminate, persistent, sparsely pu- 

 berulous on the outer surface. Leaves 5-6-jugate, the pairs 12-20 mm. apart. Pet- 

 ioles 10-15 mm. long, subalate-canaliculate; rachis 6.5-9 cm. long, the rachis 

 and petioles puberulous on the upper surface, glabrous beneath. Leaflets 33-72 

 mm. long, 12-18 mm. wide, elliptic or oblong-elliptic, densely puberulous above 

 on the costa, glabrous beneath, the venules prominulous. Inflorescences 7-9 cm. 

 long, the axis glabrous, the peduncles 2-6 mm. long, sparsely and minutely pu- 

 berulous; pedicels 5-6.5 mm. long, glabrous; bracteoles 8 mm. long, 4 mm. wide, 

 oblanceolate, glabrous externally, sparsely appressed-puberulous within. Hypan- 

 thium 4.5-5 mm. long, on a stipe 2.5 mm. long, glabrous. Sepals 7 mm. long, 2.5-3 

 mm. wide, oblong, rotund-obtuse, glabrous. Corolla and androecium unknown. 

 Stigma capitellate. Style about 15 mm. long, pilosulose basally. Ovary 2.5 mm. 

 long, 1.5 mm. wide, oblong-ob lanceolate, the margins densely pilosulose, the lat- 

 eral surfaces glabrous, 3-4-ovulate; free portion of the gynophore 3 mm. long, pi- 

 losulose. Fruit unknown. 



Type Collection: /. Triana 4419, "Istmo de S n . Pablo, Nouve lie -Grenade. - 

 Prov. de Choco, hauteur 70 metr.," March 1853 (HOLOTYPE NY, isotypes BM, 

 COL, US). Known only by the type collection. 



The only near relative of this rather complex species is certainly M.trinitense, 

 although there is a very remote connection with M. stenosipbon, M. colombianum 

 is set off from its nearest relative by its leaves having a greater number of pairs 

 of differently shaped leaflets, by its more or less pubescent ovary, and by the 

 geographic distribution. 



The varieties composing this species are based primarily on the distribution 

 of the ovary pubescence, number of pairs of leaflets per leaf, shape of the leaf- 

 lets, size and shape of the stipules, and the geographic distribution. Var. mona- 

 gasense, with its numerous pairs of lanceolate-oblong leaflets and pubescent 

 ovary, is not particularly closely related to either var. metaense or var. ocuma- 

 tense, its nearest relatives. These latter varieties also have pubescent ovaries 

 and/or fruit and differ from one another in the number of leaflet pairs, size of the 

 stipules, and geographic distribution. 



