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MEMOIRS OF THE NEW YORK BOTANICAL GARDEN [Vol. 8, No. 4 



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tion between the varieties could not be established. The specimens of the typical 

 variety which have nearly smooth trichomes are otherwise well-marked by the 

 large leaves; the roughening of the hairs is most marked in the Cuzco collections. 

 Several sheets from Cuzco have gland-tipped trichomes on the hypanthium, se- 

 pals, bracteoles, and pedicals: West 6422 p.p. (GH p.p.), Herrera 3208 p.p. (NY 

 p.p., F), and Cook & Gilbert 1171 (US). This has not been deemed worthy of any 

 formal recognition because of variation within a single collection and similar 

 glandulosity fluctuation in other species. 



In addition to the specimens cited for B. quinquenerve , Raimondi 7238 (USM) 

 from Tambillo in Dept. Ayacucho should be considered. This collection has mi- 

 nutely roughened hairs; elliptic, 3*nerved leaf blades, 12-21 x 5-7 mm.; and flow- 

 ers similar to B. quinquenerve . It may represent a distinct species or a variety of 

 B, quinquenerve. In habit, it is very suggestive of var. pusillum y except for the 

 3-nerved leaf blades and roughened hairs. The densely strigulose lower leaf sur- 

 faces, ternate flowers, and glandular-ciliate acute petals separate it from B. hu- 

 ancavelicae\ the much denser and roughened pubescence and glandular-ciliate 

 petals, from B. grisebacbii. 



2. Brachyotum campanulare (Bonpl.) Triana, Trans. Linn. Soc. 28: 48. 1871. 



Rhexia camp anul arts Bonpl. Rhexies 35. 1806-1808. . 



Arthrostemma campanulare (Bonpl.) DC. Prodr. 3: 136. 1828. 



Cbaetogastra campanularis (Bonpl.) Naudin, Ann. Sci. Nat. III. 14: 130. 1850. 



Trichomes smooth. Branchlets rounded-quadrangular, densely fine-strigose to 

 fine hirsute. Petiole 3*7 mm. Blade 13-27 x 11-14 mm., elliptic to elliptic-ovate 

 with the apex broadly acute to obtuse and the base obtuse, the 5 primaries im- 

 pressed above and elevated below, the 10-20 pairs of secondaries obscurely visi- 

 ble above but hidden by the pubescence below; above densely strigose, the hairs 

 8-18/mm. 2 with their bases sometimes on very low callosities; below very densely 

 loose-sericeous-strigose 20-35/mm. 2 Flowers constantly 4-merous, mostly crowded- 

 ternate with the dichasium subtended by leaves, rarely solitary or with an addi- 

 tional pair of flowers at the node below the dichasial node. Pedicel 0.5-2 mm. 

 below the bracteoles, 1-5 mm. above; pedicellar bracteoles 3-9 x 0.6-0.9(-4) mm., 

 linear to ovate, thin, above glabrous, below densely strigulose, mostly caducous 

 before anthesis. Hypanthium 6-7.5 x 3.5-4.5 mm., 0.2-0.3 mm. thick medianly, 

 densely loose-strigose, the fine hairs 15-20/mm. 2 Sepals 7-9 x 3"3.5 mm., nar- 

 rowly oblong, often slightly narrowed from middle to base and tapering to the 

 acute apices only in the terminal 2-4 mm., united at bases about 1 mm., the si- 

 nuses rounded-acute, inside usually sparsely strigulose on the apical Pet- 

 als deep purple, 12-18 x 9-11 mm., obovate or slightly obovate and symmetrical, 

 the apices narrowly obtuse to broadly acute, the gland-tipped cilia 0.1-0.6 mm. 

 (the terminal one 0.8 mm.). Filaments 4-5.5 mm., anthers 5-5.5 mm.; connective 

 at anther base 1-1.2 mm., free of the anther 0.5-0.6 mm., the ventral lobing 0.1- 

 0.2 mm. Style 17-25 x 0.5 mm., exserted 5-7 mm. Ovary 5 x 2.5-3 m m., densely 

 strigulose on the apical 2.5 mm., the apical lobes 1 mm. above the locules. 



Type Collection and Locality: Bonpland s.n. (HOLOTYPE presumably in Herb. 

 Humboldt & Bonpland at P; isotypes F, P); "in Peruviae frigidis, juxta Loxam 

 . . . pres de 2000 metres." 



Type Photographs and Illustrations: F36135 (presumed holotype or isotype); 

 Rhexies pi. 14 (1806-1808) (as Rhexia campanularis). 



Distribution: Prov. Loja, Ecuador, alt. 2000-3100 m. 



Between San Lucas and Ona, Hitchcock 21550 (GH, NY, US); Cordillera de Zamora 

 east of Loja, Camp E-7 1 (NY); Loja, Seemann 773.1 (K). Without province, Jameson s.n. 

 (US, W). 



