1953] 



REVISION OF BRACHYOTUM 



363 



The Paris isotype of B. campanulare cited here was used by Naudin in his 

 description of the vegetative features and type of inflorescence of Chaetogastra 

 canescens. His dissection sketch and the dissected flowers in the packet on the 

 same sheet are Rhexia canescens (Brachyotum ledifolium). These detached flow- 

 ers led to Naudin's mixed description; the leaves of B. ledifolium are never "5 

 nerviis," and the flowers are usually solitary rather than "solitariis-ternis inter- 

 dumque pluribus." 



B. campanulare is closely related to B . quinquenerve , but may be distinguished 

 by the shape of the often adaxially pubescent sepals, generally denser pubes- 

 cence, and fewer-flowered inflorescences. B. benthamianum is doubtfully distinct 

 from B. campanulare; only the sparser and coarser foliage pubescence, less acute 

 petal apices, and thicker and somewhat larger bracts differentiate the former. The 

 main veins of the bracteoles of B. campanulare vary from 1 to 5 in number, the 

 shape from linear to ovate, and the tenacity from caducous in bud to persistent un- 

 til the corolla drops. 



Another Jameson specimen s.n. (K), from (or sent from ?) Quito, is closely re- 

 lated to B. campanulare. The thin, 3-nerved, pedicellar bracteoles are ovate, 3" 

 4.5 x 1.5-2.5 mm., and caducous in bud; however the leaves are much less pubes- 

 cent above (4-6/mm. 2 ) and the calyx lobes triangular (4.8-5 x 3.7-4 mm.). 



3. Brachyotum benthamianum Triana, Trans. Linn. Soc. 28: 49. 1871. 



Trichomes smooth. Branchlets rounded-quadrangular, densely to moderately 

 short-strigose. Petiole 4-6 mm. Blade 15-24 x 7-10 mm., with length/width ratio 

 2.1-2.5, elliptic to ovate-elliptic with the apex acute and the base broadly acute 

 to narrowly obtuse, the 5 primaries impressed above and elevated below, the 15- 

 20 pairs of secondaries mostly hidden by the pubescence; above moderately long- 

 strigulose, the hairs 4-7/mm. 2 , each on a low callosity with basal %-% adherent 

 and the slightly branched base about 0.3 mm. diam.; below densely loose-strigu- 

 lose 25-30/mm. 2 , the glands solitary and obscured by the pubescence. Flowers 

 4- or 5-merous, mostly ternate with the dichasium subtended by leaves, occasion- 

 ally with an additional pair of flowers at the node below the dichasial node. Pedi- 

 cel 2-5 mm. below bracts, 1-2 mm. above. Bracts closely investing flower 2 or 4, 

 10-15 x 2-8 mm., lanceolate to ovate or elliptic with the apices acute, persistent, 

 firm, 5*nerved, outside densely strigose 9-12/mm. 2 , inside glabrous or marginally 

 sparsely strigulose. Hypanthium 6.5-7 x 4-6 mm., 0.5 mm. thick medianly, densely 

 sericeo-strigose, the hairs 8-15/mm. 2 and to 2.5—3 mm. long. Sepals 6.5-9.5 x 

 3.5-4.5 mm., oblong with acute apices, united at bases 0.8-1 mm., the sinuses 

 rounded-acute. Petals "violacea," 14-18 x 11-13 mm., obovate and slightly asym- 

 metrical with the apices obtuse to rounded, the mostly gland-tipped cilia 0.1-0.8 

 mm. (the terminal one 0.9-1.3 mm.). Filaments 4.5-5 mm.; anthers 5.5-6 mm.; con- 

 nective at anther base 1-1.3 mm., free of the anther 0.6-0.7 mm., the ventral lob- 

 ing 0.2-0.3 mm. Style 20-27 x 0.5-0.6 mm., exserted 8-10 mm. Ovary 5.5 x 3.5 

 mm., moderately strigulose on the apical 2.5-3.5 mm., the apical lobes 0.5-1.5 

 mm. above the locules. 



Type Collection and Locality: Hartweg 737 (LECTOTYPE K; isolectotypes 

 BR, G-BOIS, G-DEL, K, P, W; fragment of isolectotype F), and Seemann 773 bis 

 (syntype K); "in montibus Peruviae" (Loja, fide K sheet containing both syntypes). 



Type Photographs and Illustrations: Gleason 87-2 (isolectotype at K); F 16708 

 (destroyed isolectotype at B); Baill. Hist, des Pi. 7: 8, /. 11 (1880). 



Distribution: Prov. Loja, Ecuador, elev. 2500-3000 m. 



Above Loja, Lehmann 4922 (K); without province, Jameson s.n. (BR, US, W). 



