1953] 



REVISION OF BRACHYOTUM 



369 



nantly 5-merous), ternate or with an additional pair of flowers at the node below 

 the dichasial node, the dichasium subtended by persistent somewhat reduced 

 leaves 6-12 x 3-6 mm. Pedicel 0.5-2.5 mm. below bracts, 2-7 mm. above; pedi- 

 celiar bracts persistent or caducous, 3-7 x 1-3.5 mm., elliptic, above glabrous or 

 apically sparsely strigulose, below densely strigulose. Hypanthium 4.5-6.5 x 3.5- 

 5 mm., 0.3-0.5 mm. thick medianly, very densely long-strigulose to strigose 6- 

 15/mm. 2 Sepals 4.5-6.5 x 3"5 mm., ovate to oblong-ovate with acute to narrowly 

 obtuse and usually apicuiate apices, united at bases 0.8-1.5 mm., the sinuses 

 acute. Petals 10-18 x 8.5-12.5 mm., obovate and slightly asymmetrical with ob- 

 tuse to rounded apices, the cilia 0.1-0.5 mm. and obviously gland-tipped but the 

 glands early caducous. Filaments 4-6.5 mm., as long as the anthers; connective 

 at the anther base 1-1.6 mm., free of the anther 0.5-0.8 mm., the ventral lobing 

 0.2-0.6 mm. Style 18-28 x 0.6-0.7 mm., exserted 5-10 mm. Ovary 4.5-7.5 x 2.5-4 

 mm., moderately to densely long-strigulose on the apical 1.5-3.5 mm., the apical 

 lobes 0.6-1 mm. above the iocules. 



11a. Brachyotum maximowiczii var. maximowiczii. 

 Petiole 3-5 mm. Blade 11-20X 6-11 mm. 



Type Collection and Locality: Matbews (Fielding) 1256 (HOLOTYPE presum- 

 ably at LE; fragment of holotype BR; isotype(K); "in Peruvia ad Bajasan," Dept. 

 Amazonas. 



Distribution: Dept. Amazonas, Peru, alt. 2300-3000 m. 



Chachapoyas, Matbews 1260 (BR, K), Matbews s.n. anno 1835 (K), Mathews 45H (K), 

 Matbews 47H (K), Matbews s.n. anno 1838 (G-BOIS, K); Cerro Puma southeast of Chacha- 

 poyas, Pennell 15730 (PH); near Puente de Sigseg on Rio Sonche above Molinopampa, 

 Pennell 15786 (PH). 



lib. Brachyotum maximowiczii var. longifolium Wurdack, var. nov. 

 Petiolus 8-11 mm. Lamina 30-45 x 9-14 mm. 



Type Collection and Locality: Pennell 15857 (HOLOTYPE PH); Peru, Dept. 

 Amazonas, Cerro de Fraijaco (Huaui-Huni) northeast of Tambo de Ventilla, in dry 

 sandy soil, alt. 3000-3200 m., 7 Jul. 1948. "Shrub. Hypanthium and calyx jasper- 

 red; petals black." Known only from the type collection. 



Cogniaux described this species as 4-merous; on the holotype fragment ex- 

 amined, however, 6 of the 7 flowers were 5-merous; on the Kew isotype, all 15 

 examinable flowers were 4-merous. The leaf tubercles on the holotype fragment 

 are somewhat more prominent than usual, but the variation in development of these 

 hairs is quite great on the other collections. Among these other collections, ex- 

 cept for Matbews 47H, all of the 81 examinable flowers were 4-merous. However, 

 in Matbeus 47H, half of the 16 examinable flowers were 4-merous, half 5-merous; 

 this collection also has leaves with the length/width ratio greater than usual in 

 the typical variety, narrower more acute sepals, and smaller connective prolonga- 

 tion (0.7-0.9 mm., free of anther 0.3 mm.). Perhaps the holotype fragment and 

 Matbews 47H are hybridal variants (xB. angustifolium ?). 



Both varieties of B. maximowiczii are in appearance quite distinct from B. 

 radula, the typical variety because of the smaller leaves, var. longifolium be- 

 cause of the leaf blade length/width ratio of 2.6-3.3 [rather than 1.5-2(-2.4)L The 

 lower leaf surface reticulation is also much less obvious (or completely hidden in 

 var. longifolium) than in B. radula. 



12. Brachyotum racemosum Cogniaux, Bot. Jahrb. 42: 132. 1908. 



Trichomes smooth. Branchlets rounded-quadrangular, very densely loose- 

 slender-strigose, the hairs persistent and to 2 mm. long. Petiole 7-15 mm. Blade 

 30-75 x 10-30 mm., elliptic with the apex acute to rounded-acute and the base 



