1953] 



REVISION OF BRACHYOTUM 



373 



sanilla, Anthony & Tate 352 (US); Gualilagua near El Corazon, Acosta 7116 (F). Coto- 

 paxi: Cotopaxi, Asplund 6371 (G-DEL, S, US), Rowlee & Mixter 1127 (US); between Pilalo 

 and Zumbagua, Haugbt 2947 (NY, US). Napo-Pastaza: Papallacta, Heinrichs 601 (G-DEL). 

 Bolivar: Urcu-corral, Acosta 6616 (F). Tungurahua: near Ambato, Pacbano 178 (GH, NY, 

 US); various localities but especially near Ambato, Spruce 5147 (BR, F, G-BOIS, G-DEL, 

 GH, K, NY, P, S, W); Carihuairazo, Rorud s.n. (F); near "Paramo of Minza," Penland & 

 Summers 339 (F, NY); "Sec. Alta de Pasa," Acosta 8737 (F); near Mocha, Sodiro 468 

 (BR). Chimborazo: between Urbina and Mocha, Asplund 6926 (S); near Riobamba, Sande- 

 man 53 (K), Scbimpff 802 (A, G-DEL); Cubillm (east of Licto), Acosta 7533 (F), Acosta 

 7534 (F); between Pungala and Cusuipaccha, Scolnik 1534 (NY); Calagum near Sibambe, 

 Acosta 5483 (F); between Las Cochas and Pagma north of Huigra, Wiggins 11027 (NY). 

 Without province: eastern Cordillera, Rimbacb 9 (A, F, GH, NY, US), Rimbacb 82 (A, F), 

 Rimbacb 236 (NY, US). 



WITHOUT LOCALITY: Bonpland s.n. (fragment of presumed holotype of Cbaetogastra 

 bonplandiana F); herb. Ruiz & Pavbn s.n. (F); herb. Ventenat s.n. (possible isotype of 

 Melastoma ledifolia G-DEL). 



Vernacular Names: Sarzilejo (Bonpl. Rhexies); Puca Chaglla (Steyermark 

 52348); Illinche (Acosta 8737); Pucafichana (Acosta 8062); R umbra (Acosta 7533); 

 Zarzillejo (Dryander 2715); Puka-shakia (Quito, Lehmann 6377). 



One of the several Bonpland sheets seen from Paris was labeled "Loxa," but 

 this sheet seems to be part of the same collection as the Bonpland Purace speci- 

 mens. The herb. Ventenat specimen (G-DEL) is probably an isotype of Melastoma 

 ledifolia, since the sheet, according to the label, was given to Ventenat by La- 

 marck in whose publication Desrousseaux's description appeared. The herb. Ruiz 

 & Pavon specimen (F) was probably collected by Tafalla in Ecuador. 



From the type photograph of Melastoma ledifolia, which indicates that A. L. 

 de Jussieu's herbarium was not given to Paris until 1857, and from Naudin's ques- 

 tioning the transfer of this species to Lasiandra, it is evident that Naudin had not 

 seen the J. de Jussieu holotype of this species; if he had seen this collection, 

 the Jameson collection upon which Cbaetogastra sulphurea was based would 

 likely have been placed under Desrousseaux's species. The isotype of Chaeto- 

 gastra rostrata (F38256) in the Jussieu herbarium also was not annotated by Nau- 

 din, although he described that species from another sheet of the same Dombey 

 collection at Paris. 



In Wiggins 11027, Acosta 7534, and Haught 2947 , the ventral lobes of the con- 

 nective are free of the anther 0.1-0.5 mm., but otherwise these collections agree 

 perfectly with the vast majority of other collections. No demarcation can be made 

 between specimens with the leaves smaller and more generally ovate and the up- 

 per leaf surface densely pubescent (many Colombian collections) and those with 

 larger elliptic leaves and the upper leaf surface sparsely strigulose (many Pi- 

 chincha collections); every modification between these two mild extremes exists. 

 Rhexia canescens and Cbaetogastra bonplandiana generally belong to the former 

 category, Melastoma ledifolia and Cbaetogastra sulphurea to the latter. Lager- 

 heim (1899) noted the variability of these characters in various ecologic niches 

 on the slopes of Pichincha. 



In addition to B. gleasonii , B. ledifolium seems to be also related to B. radula 

 and B. maximowiczii; these Peruvian species differ in the petal color and shape, 

 the connective prolongation, the strictly appressed pubescence on stems and 

 lower leaf surface primaries, and the much broader-based trichomes on the upper 

 leaf surface. 



15. Brachyotum weberbaueri Cogniaux, Bot. Jahrb. 42: 133. 1908. 



Trichomes markedly shaggy. Branchlets rounded-quadrangular, densely and 

 persistently fine-hirsutulous, the hairs to 1.5 mm. long. Petiole 8-10 mm. Blade 

 17-30 x 6-13 mm., elliptic with the apex blunt-acute and the base acute, the 3 



