1953] 



REVISION OF BRACHYOTUM 



375 



the apices broadly rounded to obliquely truncate, the midvein sometimes with a 

 small apiculus, the non-glandular cilia 0„l-0.4 mm. (the terminal one 0.6-1 mm.). 

 Filaments 5-9 mm.; anthers 3o5-7.5 mm.; connective at anther base not at all ex- 

 tended nor free of the anther. Style 17-24 x 0.6-0.8 mm., exserted 2-6 mm. Ovary 

 4-5 x 2-2.5 mm., sparsely strigulose on the apical 0.3*1 mm. with conic setae, 

 the apical lobes 0.2-0.4 mm. above the iocules. 



Type Collection and Locality: Spruce 6084 (HOLOTYPE K; isotypes BR, G- 

 BOIS, G-DEL, GH, P, S, W); "in silvis montis Tunguragua alt. 10,000 ped." (fide 

 holotype), Prov. Tungurahua, Ecuador. 



Type Photographs: Gleason 87-5 (holotype); F 16710 (destroyed isotype at B). 



Distribution: central to southern Ecuador, alt. 2400-3000 m. 



Tungurahua: between "Leito y La Cima," Acosta 9075 (F). Azuay: near Sevilla de 

 Oro, Camp E-4586 (NY). Loja: "Cerros de Acacana" about 30 km. north of Loja, Espi- 

 nosa E1437 (NY). 



B. gracilescens and its relatives, B.. fraternum and B. rugosum, form a group 

 whose other relationships are obscure; they are somewhat suggestive of B. rost- 

 ratum and its relatives in the slender peduncles, obtuse to truncate petals, and 

 etuberculate anthers; however, the smooth hairs and/or large leaves are anomalous. 



17. Brachyotum fraternum Wurdack, sp. nov. 



Trichomata laevia. Ramuii novelli quadrangulati cum petiolis pedicellisque 

 modice laxo-strigulosi. Petiolus 3" 5 mm. Lamina 10-15x5-8 mm. ovata vel 

 elliptico-ovata apice acuta basi obtusa, nervis primariis 3 supra impressis subtus 

 expressis, secundariis 10-15-jugis supra invisis subtus obscure expressis et laxe 

 reticulatis; supra modice strigulosa, pilis gracilibus 4-6/mm. 2 per dimidium ba- 

 salem adhaerentibus; subtus in nervis primariis sparse strigulosa, nervis secun- 

 dariis superficieque sparsissime strigulosis, punctis nigris 4-7/mm. 2 ex glandu- 

 larum acervulis compositis. Inflorescentiae oppositae in axillis foliorum superi- 

 orum; flores 4-meri saepe terni; pedunculus gracilis efoliatus 7-15 x 0.5 mm., 

 apice bracteis duabus ascendentibus singulis 3.5-4 x 2.5-3 mm. ovato-triangulari- 

 bus supra glabris marginibus sparse appresso-ciliatis subtus per costam sparse 

 strigulosis aliter glabris; pedicelli 1-3 X 0.4-0.5 mm. ebracteolati. Hypanthium 

 4.5-5 x 4.5-5 mm., medio 0.3 mm. crassum, sparse brevi-strigulosum 7-9/mm. 2 

 Sepala 5*5.5 x 4-4.5 mm. ovata-oblonga apice acuta basi 1 mm. cohaerentia, sinu 

 acuto. Petala 13*14 x 8.5*11 mm. asymmetrice obovata apice late obtusa vel ro- 

 tunda, ciliis glandulosis (glandulis valde caducis) 0.1-0.2 mm. longis. Filamenta 

 7-8 mm.; antherae 5*6 mm.; connectivum basi antherae non productum nec liberum. 

 Stylus 19*20 x 0.5 mm., per 6-7 mm. exsertus. Ovarium 5 x 3.5 mm., apice per 2 

 mm. sparsissime brevi-strigulosum setis conicis, lobis apicalibus super loculos 

 0.4 mm. 



Type Collection and Locality: Camp E-4871 (HOLOTYPE NY); Ecuador, Azuay- 

 Oriente border, Paramo del Castillo and surrounding forested areas, crest of the 

 eastern cordillera on the trail between Sevilla de Oro and Mendez, alt. 11000- 

 11300 ft., 21 Aug. 1945. "Series of loose straggling shrubs, rarely more than 1 m. 

 high. Lvs deep green above, pale below, nitid on both sides. Hypanthium and 

 lobes pale green, often reddish-tinged above. Corolla urceolate in outline. Petals 

 white (greenish-tinged in bud and pale green along the veins at anthesis), margin 

 — sometimes to depth of 1 mm. — outlined in deep purple, this character standard 

 for all plants seen and evident even in young buds." Known only from the type 

 collection. 



B. fraternum is very closely related to B. gracilescens, but differs in the much 

 smaller leaves, shorter peduncles, broader peduncular bracts, all flowers in the 

 dichasia without pedicellar bracteoles, and the glands on the under surfaces of 



