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MEMOIRS OF THE NEW YORK BOTANICAL GARDEN [Vol. 8, No. 4 



the leaves in small clusters turning black with age. In all ways, this species 

 seems to be a much-reduced higher-altitude version of B. gracilescens. 



18. Brachyotum rugosum Wurdack, sp. nov. 



Trichomata modice muriculata. Ramuli novelli obscure quadrangulati modice 

 longo-strigulosi tarde glabrescentes. Petiolus 7-13 mm. Lamina 24-50 x 13-25 

 mm. ovata apice acuta basi obtusa vel subtruncata, rugosa, nervis primariis 5, 

 secundariis reticularis, omnibus supra anguste impressis subtus anguste expressis; 

 supra sparse strigosa (secus nervos primarios glabra), pilis 2/mm. 2 basi % ad- 

 haerentibus et leviter expansis (ad 0.2 mm. diam.); subtus in nervis primariis dense 

 strigosa, in venulis sparse laxo-brevi-strigosa, glandulis solitariis sparsis 2/mm. 2 , 

 aliter glabra. Flores 4-meri bini, inflorescentiis in axillis oppositis foliorum su- 

 periorum; pedunculi pedicellorumque bracteolae valde caducae non visae; pedi- 

 cellus 1-2.5 mm. sub bracteolis, 3-5 mm. super. Hypanthium 4 x 4.5 mm., medio 

 0.2 mm. crassum, sparse laxo-gracili-brevi-strigulosum 7-9/mm. 2 Sepala 4.5-5 x 



4- 4.5 mm. apice subulata (3 x 1.5 mm.) basi per 0.7 mm. cohaerentia lobis basi 

 expansis, sinu obtuso. Petala 11-12x9-10 mm. obovata apice late asymmetri- 

 ceque obtusa, ciliis 0.1-0.3 mm. (terminali 0.7 mm.) non-glandulosis. Filamenta 



5- 5.5 mm.; antherae 4. 5-5. 5 mm.; connectivum basi antherae 0.6-0.7 mm. minute 

 (per 0.1-0.2 mm.) liberum, lobis ventralibus 0.1-0.2 mm. Stylus 18 x 0.6-0.7 mm., 

 per 5 mm. exsertus. Ovarium 4.5 x 2.5 mm. apice per 2 mm. sparse brevi-strigu- 

 losum setis conicis, lobis apicalibus super loculos 0.9 mm. 



Type Collection and Locality: Steyermark 54316 (HOLOTYPE F; isotypes NY, 

 US); Ecuador, Prov. Santiago-Zamora, trail between Pailas and El Pan, ait. 2255- 

 3445 m., 10 Sept. 1943. "Shrub 5 ft. tall; fls nodding; petals dark purple; fila- 

 ments lavender; anthers whitish; leaves rugose both sides, dark green above, dull 

 green below." Known only from the type collection. 



B. rugosum is more suggestive of the B. rostratum complex than either of its 

 relatives, but differs considerably in its large rather thin leaves, eglandular petal 

 cilia, and much less prominently roughened hairs. From its nearest relatives, B. 

 gracilescens and B. fratemum, it differs in the roughened hairs, 5-nerved leaves, 

 narrower calyx lobe apices, and longer ovary lobes. The flower dissected was 

 from the New York isotype. 



19. Brachyotum microdon (Naudin) Triana, Trans. Linn. Soc. 28: 49. 1871. 



Chaetogastra microdon Naudin, Ann. Sci. Nat. III. 14: 132. 1850. 

 Chaetogastra pentlandii Naudin, Ann. Sci. Nat. III. 14: 133. 1850. 

 Chaetogastra hermannioides Naudin, Ann. Sci. Nat. III. 14: 133. 1850. 

 Brachyotum pentlandii (Naudin) Triana, Trans. Linn. Soc. 28: 49. 1871. 

 Brachyotum hermannioides (Naudin) Triana, Trans. Linn. Soc. 28: 49« 1871. 

 Brachyotum setosum Gleason, Mem. N. Y. Bot. Gard. 7: 314. 1927. 



Tri<?homes minutely but densely roughened. Branchlets quadrangular, sparsely 

 to densely strigulose to loose-strigulose. Petiole 4-20 mm. Blade 15-85 x 8-40 

 mm., elliptic or elliptic-lanceolate to ovate with the apex acute to rounded and 

 the base obtuse to subtruncate, thinly coriaceous to chartaceous, the 3 primaries 

 thinly impressed above and elevated below, an additional pair of marginals more 

 or less distinctly developed, the numerous soon-reticulate secondaries obscure 

 above and obscurely elevated below; above sparsely to moderately strigulose or 

 strigose, the hairs 1-15/mm. 2 with the basal %-% adherent and not abruptly ex- 

 panded; below sparsely to moderately hirsute or strigulose (l-)2-15(-25)/mm. 2 , 

 the glands solitary and sparse to very dense 5-80/mm. 2 Flowers constantly 5" 

 merous, mostly ternate, sometimes with a pair of additional flowers at the node 

 below the dichasial node or with still another pair at the next-lower node, occa- 

 sionally the dichasia ternate, the dichasia subtended by persistent slightly re- 



