378 



MEMOIRS OF THE NEW YORK BOTANICAL GARDEN 



[Vol. 8, No. 4 



lobes 1-1.5 mm. long are Balls B6249 p.p. (US p.p.), Steinbach 8667,Herzog 7 923, 

 and Cuming s.n. With similar leaf shape but denser and longer upper leaf surface 

 pubescence (6-14/mm. 2 ) and very loosely appressed dense branchlet pubescence 

 are Werdermann 2015, Steinbach 8512, and most of Balls B6249, although the last- 

 named collection has great variation in pubescence density. Balls B6277 ranges 

 from sprigs similar to Balls B6249 to some approaching the northern "herman- 

 nioides" element (but with contracted inflorescences); also similar to the north- 

 ern "hermannioides" phase is Steinbach 5948. In part overlapping these last two 

 collections to some extent is a "microdon" element with compact inflorescences, 

 elliptic-lanceolate acute-tipped leaf blades with length/width ratio of 2-3.6 and 

 with the upper surfaces moderately strigulose ((3-)5-8(-ll)/mro. 2 ), generally very 

 loosely appressed branchlet pubescence, and ovary apex lobes 1.5-2.5 mm. long. 

 This group includes all other specimens from Depts. La Paz and Cochabamba, ex- 

 cept Cardenas 3056 which has a leaf blade length/width ratio of 2.6-3 but ovary 

 apex lobes 3.7-3.9 mm. long. In all the foregoing groups, the pedicellar bracteoles 

 are mostly caducous before anthesis and generally are linear to very narrowly 

 spatulate (5*14 x 0.7-4 mm.); the pedicels are 2-7(-ll) mm. long below the brac- 

 teoles. With leaf shape and size as in Balls B-6277 p.p. and Steinbach 5948 

 (ovate and acute-tipped, 20-40 x 10-20 mm.), but branchlet pubescence sparse 

 and strictly appressed, the upper surfaces of the leaf blades sparsely short- 

 strigose (1-3/mm. 1 ), the pedicels 10-20 mm. long below the bracteoles, and the 

 persistent bracteoles 8-11 x 1.5-3.5 mm., is a northern "hermannioides" element: 

 Weddell 3783, Weddell 3784, and Carriker s.n. The southern "hermannioides" 

 element, consisting of Fiebrig 2458, Fries 1283, and all Argentinian specimens, 

 is similar but shows larger leaves (30-85 x 20-40 mm.), variation in length of 

 pedicels below bracteoles of 6-60 mm., and persistent bracteoles 10-41 x 2-11 

 mm. While the morphologic variation between the extremes is great, even varietal 

 delimitation within B. microdon will be difficult and has been deferred. 



No immediate relatives of B. microdon are apparent. The pubescence is some- 

 what suggestive of B. ledifolium. Perhaps a closer relative is B. floribundum, 

 which differs in type of pubescence, development of inflorescence, and size of 

 flowers. 



20. Brachyotum sanguinolentum (Naudin) Triana, Trans. Linn. Soc. 28: 49. 1871. 



Chaetogastra sanguinolenta Naudin, Ann. Sci. Nat. III. 14: 131. 1850. 

 Brachyotum floribundum (GrisebacM Triana, Trans. Linn. Soc. 28: 49. 1871. Nomen. 

 Brachyotum floribundum (Grisebach) Triana ex Cogniaux; DC. Monog. Phan. 7: 155. 

 1891. 



Brachyotum barbiferum Rusby, Phytologia 1: 69. 1934. 



Brachyotum pedicellatum Markgraf, Notizbl. Bot. Gart. Berlin 13: 460. 1937. 



Tricjiomes smooth. Branchlets notably quadrangular, nearly glabrous to moder- 

 ately short-strigose or the pubescence spreading. Petiole 2-18 mm. Blade 17-76 x 

 9-25 mm., elliptic with the apex acute to rounded-acute or obtuse and the base 

 acute to obtuse, the 3 primaries (occasionally an additional pair of marginals 

 varyingly developed) and 15-30 pairs of secondaries obscurely to noticeably (but 

 narrowly) impressed above and finely elevated below, the secondaries usually re- 

 ticulate; above glabrous or very sparsely strigulose to moderately short-strigose, 

 the hairs to 7/mm. 2 with their basal adherent; below nearly glabrous to mod- 



erately loose-short-strigose or hirsutulous, the hairs to 15/mm. 2 , the glands usu- 

 ally solitary and 6-20/mm. 2 -but occasionally in clumps. Flowers constantly 4- 

 merous, in terminal corymbiform usually ternate few- to many-flowered cymes, the 

 cymes unbranched to twice-branched below the cymes, the ultimate pedicels soli- 

 tary to ternateiy dichasial. Pedicel 0-10 x 0.4-0.5 mm. below the bracteoles, 3- 



