380 MEMOIRS OF THE NEW YORK BOTANICAL GARDEN [Vol. 8, No. 4 



I 



specimens would correspond to B. floribundum, including B. barbiferum. Pearce 

 s.n. has the large leaves of this latter group, the vegetative pubescence density 

 of the "sanguinolentum" element, and hypanthial pubescence density of 2/mm. 2 

 Weberbauer 626 has the branchlet, peduncle, and pedicel pubescence spreading, 

 the blade apices often rounded, and the lower leaf surface pubescence denser than 

 in the "floribundum" element, but otherwise agrees in leaf size and pubescence 

 density with that element. Such variation in degree of appression of pubescence 

 has been noted in other species (B. grisebachii, B. rostratum) where intermediate 

 degrees were fully represented, and does not seem worthy of formal recognition. 

 Cogniaux's limited citations of dimensions suggest an inverse quantitative corre- 

 lation between leaf and flower size, but such a relation does not exist. Success- 

 ful infraspecific differentiation based on leaf size and pubescence density seems 

 probable, but has been deferred. 



The closest relative of B. sanguinolentum would appear to be B. grisebachii, 

 which differs greatly in inflorescence development, calyx lobes, and petal shape. 

 The only species with a similar degree of inflorescence development is B. 

 quinquenerv e, 



21. Brachyotum grisebachii Cogniaux; DC. Monog. Phan. 7: 153* 1891. 



Trichomes smooth. Branchlets quadrangular, very sparsely hirsute to strigu- 

 lose, very soon glabrescent. Petiole 2-4(-7) mm. Blade 8-24 x 5—11 mm., ovate- 

 lanceolate to lance-elliptic or elliptic to ovate with the apex acute or rounded- 

 acute and the base obtuse to subtruncate, the 3 primaries impressed above and 

 elevated below, the 7-12 pairs of secondaries narrowly to obscurely impressed 

 above and obscurely elevated below, the margins i, s. usually strongly recurved; 

 above glabrous (sparsely strigulose when young) or with some marginal hairs per- 

 sisting or sparsely and permanently strigulose 1(— 2)/ 1—2 mm. 2 ; below sparsely 

 hirsutulous to sparsely rtrigulose 1-4/mm. 2 , the glands in clusters often forming 

 minute black punctae with age. Flowers constantly 4-merous, mostly crowded- 

 ternate, sometimes with an additional pair at the node below the dichasial node, 

 rarely solitary; dichasial peduncle erect, not differentiated from the branchlets, 

 the internode below the dichasium 5-11 x 1.1-1.7 mm., the dichasium subtended 

 by slightly reduced and persistent leaves. Pedicel 0.5-2 mm. below bracteoles, 

 2-6 mm. above; pedicellar bracteoles leaflike to noticeably modified, 2-10 x 1-2.5 

 mm., oblong to obovate, persistent or caducous before anthesis, above glabrous or 

 apically very sparsely strigulose, below very sparsely strigulose. Hypanthium 4- 

 7 x 3.5-5 mm., 0.3 mm. thick medianly, moderately strigulose with the thin hairs 

 (4-)8-12/mm. 2 Sepals (4.5-)6-10(-l 1.5) x'(3.5-)4-4.5 mm., lanceolate with nar- 

 rowly acute apices, united at bases 0.7-1.3 mm., the sinuses acute. Petals deep 

 purple, ^13-^0 x 9-15 mm., obovate to elliptic and asymmetrical with the apices 

 acute to narrowly obtuse, occasionally outside very sparsely strigulose basally, 

 the marginal cilia 0.2-0.5 mm. (the terminal few to 1 mm.) and eglandular (very 

 rarely with rather persistent glandular tips). Filaments 5-6.5 mm.; anthers 4-8 

 mm.; connective at anther base 1-1.5 mm., free of the anther 0.6-0.7 mm., the 

 blunt ventral lobes (0.1-)0.4-0.7 mm. Style 17-26 x 0.4-0.5 mm., exserted 5-8 mm. 

 Ovary 4.5-6 x 2.5-4 mm., moderately strigulose on the apical 1.5-3 mm., the ap- 

 ical lobes 0.6-0. 7(-l. 4) mm. above the locules. 



Type Collection and Locality: Lechler 1856 (LECTOTYPE collection BR, 

 G-BOIS, K, P, S, W; fragment. F); Peru, Dept. Puno, "prope Agapata." 



Type Photographs: Gleason 51-2 (isolectotype, presumably in G-DC); F 16711 

 (destroyed isolectotype at B). 



Distribution: southeastern Peru to northwestern Bolivia, alt. 3000-3700 m. 



