382 MEMOIRS OF THE NEW YORK BOTANICAL GARDEN [Vol. 8, No. 4 



cavelicae are often so crowded on the densely leafy branchlets that their solitary 

 disposition is difficult to observe; yet in all specimens, terminal vegetative 

 growth of the floriferous branches can be seen. The leaves on herbarium material 

 show the same deeply impressed veins on the upper surfaces as found in B.grise- 

 bachii and B. tyrianthinum. From B . ty riant hinum, B. huancavelicae may be distin- 

 guished by the smooth or only very minutely and sparsely roughened hairs, the 

 very narrowly acute calyx lobes which are somewhat expanded basally and gla- 

 brous within, and the non-glandular petal cilia. From B. naudinii, it differs in the 

 generally larger leaves and sepal shape, as well as the non-glandular petals. B. 

 grisebachii may be separated by the ternate flowers and generally less dense pu- 

 bescence, as well as the more acute petals. Vargas 319 (F), from Paucartambo in 

 Dept. Cuzco, has the pubescence density of B. huancavelicae, but ternate flowers 

 with the petals basally pubescent outside; Macbride (1941, p. 274) referred this 

 specimen to B. strigosum, a quite different Colombian species; this imperfect 

 collection is here doubtfully referred to B. grisebachii. 



23. Brachyotum nutans Gleason, Bull. Torrey Club 54: 25. 1927. 



Trichomes smooth. Branchlets quadrangular, very sparsely strigulose and soon 

 glabrescent. Petiole 0.5*3.5 mm. Blade 5*12 x 3*6.5 mm., ovate to elliptic-ovate 

 with the apex rounded and the base obtuse to subtruncate, the 3 primaries nar- 

 rowly expressed above and elevated below, the secondaries obsolete; above gla- 

 brous; below very sparsely strigulose on the primaries, glabrous on the surface 

 except for a few tufts of glands which turn black with age. Flowers constantly 4- 

 merous, solitary on short lateral branchlets in opposite upper leaf axils, the 

 branchlets with 1-2 pairs of leaves. Pedicel 2*5 mm. above the last slightly- 

 reduced leaves (bracteoles ?). Hypanthium 3.5*4.5 x 3.5*4 mm., 0.2-0.3 mm. thick 

 medianly, very sparsely strigulose (1/1-3 mm. 2 ) or glabrous except for a few api- 

 cal hairs. Sepals 3.5*5.5 x 2.5*4.5 mm., triangular with acute apices, outside 

 glabrous except for a very few appressed hairs basally, united at bases 0.5*1 mm., 

 the sinuses acute. Petals deep purple, 15*18 x 11-14 mm,, asymmetrically obo- 

 vate with the apices bluntly obtuse, the gland-tipped cilia 0.1-0.2 mm. Filaments 

 3.5*5.5 mm.; anthers 4*5 mm.; connective at anther base 0.7-1.3 mm., free of the 

 anther 0.4-0.7 mm., the ventral lobing 0.1-0.4 mm. Style 17-27x0.3-0.5 mm., 

 exserted 6-11 mm. Ovary 4.5*5 x 2*3.5 mm., moderately to densely strigulose on 

 the apical 1-2.5 mm., the apical lobes 0.3-0.6 mm. 



Type Collection and Locality: Pennell 13847 (HOLOTYPE NY; isotypes F, 

 GH, PH, S, US); Peru, Dept. Cuzco, Paso de Tres Cruces. 



Type Photograph: New York s.n. (holotype). 



Distribution: Dept. Cuzco, Peru, alt. 2800-4000 m. 



Pjnasniocc, Cook & Gilbert 1244 (US); Ollantaitambo, Herrera 3342a (US); "Acanacu" 

 near Paucartambo, Balls B6714 (NY), Vargas 320 (F), West 7034 (GH); Paucartambo, 

 Soukup 379 (F); Marcapata, Stafford 990 (K). 



B. nutans may be little more than a depauperate variation of B. naudinii, but 

 the nearly glabrous leaves and hypanthia hold the cited specimens in a compact 

 unit. The acute sepals are more like those of B. grisebachii which, however, has 

 larger leaves, ternate flowers, much more densely pubescent hypanthia, and pet- 

 als of a slightly different shape. 



24. Brachyotum naudinii Triana, Trans. Linn. Soc. 28: 48. 1871. 



Trichomes smooth to very minutely and sparsely roughened. Branchlets quad- 

 rangular, moderately to sparsely short-strigulose. Petiole 2-5 mm. Blade 5-17 x 

 4-7(-10) mm., ovate to elliptic with the apex rounded to blunt-acute and the base 



