1953] 



REVISION OF BRACHYOTUM 



383 



obtuse to subtruncate, the 3 primaries narrowly impressed above and elevated be- 

 low, the secondaries obscure or obsolete above and below; above sparsely strigu- 

 lose, the hairs 1-4/mm. 2 with their basal %-% adherent; below sparsely strigu- 

 lose on the primaries, very sparsely on the surface 2-6(-10)/mm. 2 , the glands to 

 25/mm. 2 and solitary or in small clusters at hair bases. Flowers usually solitary 

 on short lateral branchlets, usually exclusively 4-merous, occasionally predom- 

 inantly 5~merous. Pedicel 2-5 rnm. above the last somewhat reduced persistent 

 leaves (bracteoles ?). Hypanthium 4-7 x 3.5-6 mm., 0.3-0.5 mm. thick medianly, 

 sparsely to moderately strigulose, the fine hairs (6-)8-l6/mm. 2 Sepals (4-)5-7 

 (-9) x (3-)4-5 mm., oblong-ovate (rarely deltoid-ovate) with the apices rounded to 

 broadly acute, united at bases 0.7-1.1 mm., the sinuses rounded-acute. Petals 

 deep purple, 13-23 x 10-17 mm., asymmetrically or nearly symmetrically obovate 

 with the apices broadly obtuse to rounded or truncate, the gland-tipped cilia 0.1- 

 0.4(-0.6) mm. Filaments 3.5-7.5 mm., as long as the anthers; connective at anther 

 base (0.6-)l-1.6 mm., free of the anther for Vi the length or slightly less, the 

 blunt ventral lobes 0.2-0.4(-0.8) mm. Style 17-25 x 0.4-0.6 mm., exserted 4-7 

 mm. Ovary 4.5-7 x 3-4 mm., sparsely to moderately strigulose on the apical 

 (0.6-)1.5-3 mm., the apical lobes 0.3-1 mm. above the locules. 



Type Collection and Locality: Dombey s.n. (HOLOTYPE presumably at P; 

 isotypes BM, BR, F, G-DEL, L, P); "Peruvia," Dept. Huanuco (or more probably 

 Junln), Palca, fide Macbride (1941, p. 271). 



Type Photograph: F36134 (presumed holotype at P). 



Distribution: central (to northern ?) to southeastern Peru, alt. 2700-4000 m. 



Ancash: Banos de Chancos near Huaras, Sandeman 4610 (K). Junm: east of Huancayo, 

 Stork & Horton 10218 (F, UC). Cuzco: Ollantaitambo, Cook & Gilbert 720 (US); Yucay, 

 Soukup 736 (F, GH); Hda. Araypallpa near Paruro, Vargas 389 (GH); "Valle de Apuri- 

 mac," Herrera s.n. (F). Ayacucho: Quinua, Weberbauer 5540 (F); "Totorobamba" in Prov. 

 Huamanga, Weberbauer 5484 (F, GH); "Pallcea" between "Pajanal" and Ayacucho, Balls 

 B6928 (GH, NY, UC, US); south of "Puchuhuillca" near "Mataral," West 3662 (GH, UC). 

 Apunmac: north of Chincheros, Stork & Horton 10763 (F, UC); km. 58 between Abancay 

 and Ayacucho, Ferreyra 2795 (NY); "Quisvala" northeast of Abancay, Bills B6894 (GH, 

 NY, UC, US). Without Department: "Chile," Gay s.n. (F); "Lima," Gay s.n. (P); "Pace- 

 chac," Hill 153 (K). 



Vernacular Names: Masuca (Cook & Gilbert 720); Ccehuincha (West 3662); 

 Jehuincha (Stork & Horton 10763). 



One Paris isotype is labeled only "herb. Richard"; this specimen was used 

 by Cogniaux in drawing up his erroneous description of B. microphyllum; the iso- 

 type in Cogniaux's herbarium (BR) is a branch from this Paris sheet; while no 

 collector was indicated on the Paris specimen, it is undoubtedly part of the Dom- 

 bey type coilection. 



In the type collection, 17 of the 28 examinable flowers were 5-merous; in 

 Sandeman 4610, 9 of 9; and in Stork & Horton 10218 t 8 of 10. The remainder of 

 the flowers in these collections and all of the 82 examinable flowers in all other 

 cited specimens were 4-merous. The degree of pubescence roughening varied in- 

 dependently of the mery, the first two collections with predominantly 5"merous 

 flowers being perceptibly roughened, and the third-mentioned smooth. The con- 

 stantly 4-merous collections have pubescence scabridity between these extremes. 



B. naudinii may be distinguished from B. grisebacbii by the shape of the se- 

 pals and petals, the glandular petal cilia, and the solitary flowers. From B. tyri- 

 anthinum, it differs in the smaller leaves and shorter, sparser, less roughened 

 trichomes, as well as the adaxially glabrous sepals. However, there are a series 

 of constantly 4-merous specimens from northern Peru having pubescence develop- 



