384 MEMOIRS OF THE NEW YORK BOTANICAL GARDEN [Vol 8, No. 4 



ment very suggestive of B. tyrianthinum, but the trichomes are smooth; the pubes- 

 cence on the upper leaf surface ranges from 3-9/mm. 2 and the sepals are rather 

 variable in shape, ovate to ovate-oblong with variably acute apices, but glabrous 

 within. They include Stork & Horton 9962 (F, G-DEL, UC) from La Libertad, Fer- 

 reyra 3291 (USM) and Raimondi 3152 (USM) from Cajamarca, and Mathews 1259 

 (K) and Pennell 15530 (PH) from Amazonas. These specimens have been doubt- 

 fully referred to B. naudinii. Sandeman 4291 (K) from Piura is even more anoma- 

 lous, possibly being a depauperate form of the preceding group; the calyx lobes 

 are very narrowly oblong-lanceolate and leaves narrowly elliptic. 



25. Brachyotum tyrianthinum Macbride, Field Mus. Pub. Bot. 4: 174. 1929. 



Trichomes moderately but minutely scabrid. Branchlets obscurely quadrangu- 

 lar, moderately to densely long-strigulose to strigose, tardily glabrescent. Peti- 

 ole (1— )3— 6 mm. Blade 10-30 x 6-14 mm., narrowly ovate to elliptic with the apex 

 acute or blunt-acute and the base broadly acute to subtruncate, the 3 primaries 

 narrowly impressed above and elevated below, the 8-12 pairs of secondaries 

 mostly invisible above and lightly elevated below; above moderately short-stri- 

 gose, the hairs (3-)5-7(-10)/mm. 2 with their basal adherent and not abruptly 

 expanded; below densely long-strigulose on the primaries, moderately to sparsely 

 hirsutulous or loose-strigulose on the surface (3-)7-l l/mm. 2 , the glands mostly 

 solitary 12-30/mm. 2 Flowers 4-5~merous, the 4-merous ones usually predomi- 

 nant, solitary on short lateral leafy branchlets in opposite upper leaf axils. Pedi- 

 cel 2-5 mm. above the last somewhat reduced (3-10 x 1-4 mm.) persistent leaves 

 (bracteoles ?). Hypanthium 4.5-6 x 4-5 mm., 0.3-0.5 mm. thick medianly, sparsely 

 to moderately strigose, the fine hairs 8-ll/mm. 2 and to 1.5-3.5x0.2-0.3 mm. 

 Sepals 6.5-9 x 3—4.5 mm., oblong-ovate to lanceolate with the apices acute, in- 

 side sparsely strigulose on the apical l >£-%. Petals deep purple, 12-17x9-14 

 mm., obovate and sometimes slightly asymmetrical with the apices very broadly 

 obtuse, the gland-tipped cilia 0.1-0.2 mm. (the terminal few to 0.4-0.8 mm.). Fila- 

 ments 4-5 mm.; anthers 4.5-6.5 mm.; connective at anther base 1-1.7 mm., free 

 of the anther 0.3-0.7 mm., the ventral lobing 0.1-0.2 mm.- Style 18-21 x 0.4-0.7 

 mm., exserted 5-6 mm. Ovary 6.5-7 x 3.4-4 mm., sparsely to moderately long- 

 strigulose on the apical 2-3.5 mm., the apical lobes 0.7-1 mm. above the locules. 



Type Collection and Locality: Macbride & Featherstone 1438 (HOLOTYPE F; 

 isotypes G-DEL, S, US); Peru, Dept. Huanuco, Mito, alt. 2750 m. 



Type Photograph: F63425 (holotype). 



Distribution: central to northern Peru, alt. 2750-4000 m. 



Cajamarca: between Cajamarca and Celendin, Scolnik 1323 (NY). Huanuco: Chaglla, 

 Sandeman 5109 (K); Mito, Macbride & Featherstone 2092 (F, NY); probably near Cerro de 

 Pasco, Sawada P89 (F). Without Locality: "Columbia," Lobb s,n. (K). 



Vernacular Names: Cachis (Macbride & Featherstone 1438). 



All of the 17 examinable flowers in the type collection were 4-merous, as were 

 all 15 in Scolnik 1323; 1 of 10 flowers in Sandeman 5109 > 2 of 6 in Lobb s.n. t 9 

 of 15 in Macbride & Featherstone 2092, and 5 of 5 in Sawada P89 were 5-merous. 

 Aside from mery, the species is quite uniform morphologically, although an oc- 

 casional flower has the sepals glabrous within. 



Aside from its immediate relatives, B. tyrianthinum may be related to B. ros- 

 marinifolium t the roughened pubescence being a possible indication of such a 

 linkage. It is also somewhat reminiscent of B. strigosum, which however has 

 smooth hairs, differently shaped sepals, much stouter hypanthial pubescence, 

 larger and more deeply divided connective prolongation, and smaller apical ovary 

 lobes. 



