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MEMOIRS OF THE NEW YORK BOTANICAL GARDEN [Vol. 8, No. 4 



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phologic intermediate between these two relatives should be through such gla- 

 brous large-leaved specimens of B. lindenii as Hartweg 1003 p.p.; however no 

 such intermediates have been seen. Cuatrecasas 20039, from Paramo de Bavaya 

 in Dept. Valle in Colombia (F, NY), is, however, intermediate between the two 

 species to some extent, having the leaf and hypanthial pubescence as well as the 

 inflorescences and inconspicuous nodal setae of "typical" B. lindenii and the 

 larger sepals (10-10.5 x 3.5-4 mm.) and more pointed petals of B. cernuum; it is 

 perhaps best regarded as an exceptionally robust specimen of B. lindenii. 



27. Brachyotum lindenii Cogniaux; DC. Monog. Phan. 7: 159. 1891. 



Brachyotum riveti Danguy & Chermezon, Bull. Mus. Hist. Nat. [Paris] 28: 433. 1922. 

 Brachyotum strigosum vat. tolimensis Cuatrecasas, Trab. Mus. Nac. Ci. Nat. Madrid 

 33: 89. 1936. 



Trichomes smooth, slender. Branchlets quadrangular, sparsely to moderately 

 strigulose, soon glabrescent. Petiole 1-5 mm. Blade 9-22x4-9 mm., narrowly 

 ovate with the apex acute to blunt-acute and the base obtuse to subtruncate, the 

 3 primaries narrowly impressed above and elevated below, the 6-12 pairs of sec- 

 ondaries obscurely impressed or invisible above and very lightly elevated below; 

 above glabrous to sparsely strigulose, the hairs to 5*6/mm. 2 and with the basal 

 %—% adherent; below sparsely strigulose on the primaries, the surface glabrous to 

 sparsely loose-strigulose, the hairs to 8/mm. 2 , a cluster of glands at the base of 

 each hair turning black with age. Flowers predominantly 5~merous, sometimes ter- 

 nate but mostly with an additional pair at the node below the dichasial node, oc- 

 casionally the dichasia ternate. Peduncle, at anthesis of the terminal dichasium, 

 with the internode below the dichasium 6-15 x 0.5-1 mm. and strongly pendant; 

 bracts at base of dichasium 3.5*7 x 1-3 mm., spatulate to leaf-like, in the 5~flow- 

 ered inflorescences early caducous. Pedicel 3-10 mm. below bracteoles, 1-3 mm. 

 above; pedicellar bracteoles 1-6 x 0.3-1 mm., linear to narrowly spatulate, very 

 early caducous, above glabrous, below very sparsely strigulose. Hypanthium 4* 

 6 x 3-5 mm., 0.3-0.5 mm. thick medianly, sparsely to moderately strigulose, the 

 hairs (2-)5*10(-20)/mm. 2 Sepals 4-8.5 x 2.5-4.5 mm., lanceolate with acute api- 

 ces, united at bases o 6-l.l mm., the sinuses acute. Petals deep purple, 10-16 x 

 8-13 mm., obovate and slightly asymmetrical with the apices broadly obtuse to 

 rounded, the non-glandular cilia 0.1-0.5 mm. (the terminal few 0.4-1 mm.). Fila- 

 ments 4-6 mm.; anthers 4-6 mm.; connective at anther base (0.7-)l-1.7 mm., free 

 of the anther 0.5-0.8 mm., the ventral lobing 0.3-0.7 mm. Style 15-22 x 0.5-0.7 

 mm., exserted 3-8 mm. Ovary 4.5*5.5 x 2.5-4 mm., sparsely to moderately strigu- 

 lose on the apical 1.5*2.5 mm. with the hairs usually conic, the apical lobes 0.3* 

 0.7 mm. above the locules. 



Type Collection and Locality: Linden 925 (LECTOTYPE G-DEL; isolecto- 

 types BR, K, P, W; fragments F); Colombia, Dept. Tolima, "prov. Mariquita ad 

 Tolima," "hauteur 2000 toises." 



Type Photographs: F16712 (destroyed syntype at B, collected by Stubel); 

 F36922 (syntype of B. riveti at P, Rivet 342). 



Distribution: north central Colombia to northern Ecuador, elev. 3000-4200 m. 



COLOMBIA: Antioquia: Paramo Monro Frontino north of Urrao, Core 377 (NA, NY). 

 Caldas: Manizales, Sandeman 5709 (K); Paramo del Quindio, Pennell & Hazen 9990 (GH, 

 NY, PH, US), Triana s.n. (isosyntypes of B. riveti K, P, US, W). Cundinamarca: Bogota, 

 Triana s.n. (K, NY, P, W). Tolima: Nevada del Tolima, Cuatrecasas 2834 (isosyntype of 

 B. strigosum var. tolimensis K). Cauca: Volcan del Purace, Cuatrecasas 14700 (F, NY), 

 Pennell & Killip 6567 (GH, NY, PH, US), von Sneidern 1833 (S), von Sneidem 1834 (S). 

 Narino: between Buesaco and Pasto, Andre 817 (K); Volcan El Galeras, Ewan 16330 (NY), 

 Schultes & Villarreal 7952 (NY); Pasto, Hartweg 1003 p.p. (BR, G-BOIS, G-DEL, K), 

 Andre 1075 (K); Yacuanquer, de Garganta 476 (F); Paramo del Angel, Sandeman 110 (K). 



