2 953] 



REVISION OF BRACHYOTUM 



387 



ECUADOR: Carchi: Paramo del Angel, Acosta 10546 (F), Asplund 10393 (S); "La 

 Rinconada" between Ibarra and Tulcan, Hitchcock 20792 (GH, NY, US); Nudo de Boliche, 

 Penland & Summers 866 (F, NY); San Gabriel, Holmgren 7492 (S); between "Moran and 

 Olivos," Mexia 7492 (F, NY, UC, US). Imbabura: between Ibarra and Mariano Acosta, 

 Drew E-289 (NA, NY). Pichincha: Mojanda, Sodiro 467 (BR); between Pedregal and "Hda. 

 Yanurcu," Acosta 8299 (F). 



In the packet of Sodiro 467 (BR) is B. alpinum, doubtfully from the same col- 

 lection; this sprig may be part of Sodiro 466. 



As has been noted in other species, the density of foliar pubescence is usu- 

 ally of no specific reliability, and in the case of B. lindenii and B. riveti, also of 

 no varietal significance. All intergradations between B. lindenii, with leaves 

 sparsely strigulose above and sparsely Ioose-strigulose below on the surface 

 (Triana s.n. Bogota, Linden 925, Andre 817, Core 377, Andre 1075, Ewan 16330), 

 and B. riveti, with leaves glabrous above and very sparsely strigulose only on 

 the primaries below (Sandeman 111, Mexia 7492, Hitchcock 20792, Acosta 10546, 

 Penland & Summers 866, de Gargantua 476, Schultes & Villarreal 7952, Sodiro 

 467, Drew E-289, Asplund 10393, Holmgren 878), can be found, even within some 

 of the collections cited above. Of the 156 observable flowers among all the col- 

 lections, only 15 were 4-merous (2 of 24 in Pennell & Hazen 9990,7 of 23 in Pen- 

 nell & Killip 6567, 1 of 6 in Schultes & Villarreal 7952, 1 of 9 in Sandeman 1 10, 

 1 of 3 in Acosta 10546, 2 of 8 in Hitchcock 20792, and 1 of 11 in Penland & 

 Summers 866). 



28. Brachyotum alpinum Cogniaux; DC. Monog. Phan. 7: 167. 1891. 



Trichomes smooth, slender. Branchlets quadrangular, sparsely strigulose, 

 soon glabrescent. Petiole 1-4 mm. Blade (6-)10-17x 4-9 mm., the shape and ven- 

 ation as in B. lindenii; above glabrous to very sparsely strigulose, the hairs to 

 1-2/mm. 2 and with their basal % adherent; below sparsely strigulose on the pri- 

 maries, the surface glabrous to sparsely loose-strigulose, the hairs to 7/mm. 2 , 

 the glands as in B. lindenii. Flowers constantly 4-merous, the inflorescences as 

 in B. lindenii. Hypanthium 4-7 x 4-5 mm., 0.3-0.4 mm. thick medianly, sparsely 

 strigulose, the- hairs 4-8/mm. 2 Sepals 3.5*8 x 3-4 mm., lanceolate to oblong- 

 lanceolate with acute apices, united at bases 0.5-1.2 mm., the sinuses acute to 

 rounded-acute. Petals deep purple, 10-14x8-11 mm., asymmetrically obovate 

 with the apices rounded to broadly obtuse, the eglandular cilia 0.1-0.6 mm. Fila- 

 ments 4-4.5 mm.; anthers 4.5-6.5 mm.; connective at anther base 1-1.5 mm., free 

 of the anther 0.4-0.8 mm., the ventral lobing 0.3-0.6 mm. Style 17-22 x 0.4-0.6 

 mm., exserted 3-6 mm. Ovary 4-5.5 x 2.5-3.5 mm., moderately strigulose on the 

 apical 1.5-2.5 mm., the apical lobes 0.3-1 mm. above the locules. 



Type Collection and Locality: Jameson s.n. (or 293 ?) (SYNTYPES presuma- 

 bly in G-DC and LE; isosyntypes BR, F, G-BOIS, G-DEL, K, US, W) and Eraser 

 s.n. (SYNTYPE presumably in G-DC); Ecuador, Prov. Chimborazo, "mont. Chim- 

 borazo altit. 4000 m.," and "Ecuador," respectively. 



Type Photographs: Gleason 51-5 (Jameson and Fraser syntypes presumably in 

 G-DC); F25862 (Fraser syntype presumably in G-DC). 



Distribution: north central Ecuador, alt. 3200-4000 m. 



Pichincha: "Quitensian Andes," Couthouy s.n. (GH, NY). Bolivar: "Gualicon Loma," 

 Acosta 6278 (F). Tungurahua: "Paramo of Minza," Penland & Summers 325 (F, NY). 

 Chimborazo: Chimborazo, Sodiro 466 (BR); "Chimborazo and Cayambe," Hall 3 (K); Cerro 

 Altar, Heinrichs 877 (G-DEL, NY); near Riobamba, Rimbach 137 (NY, US); between Huam- 

 boya and Pungala, Scolnik 1542 (NY); "Sinche" (Sinchan ?), Tate 461 (US); trail to "El 

 Placer," Acosta 7231 (F). Without Province: "Eastern Cordillera," Rimbach 21 (F, GH); 

 "Guyaquil," herb. Ruiz & Pavon s.n. (F). 



