1953] 



REVISION OF BRACHYOTUM 



391 



Bolina conferta (Bonpl.) Raf. Syl. Tell. 101. 1838. 



Bracbyotum campy lanthum Triana, Trans. Linn. Soc. 28: 49« 1871. 



Trichomes smooth. Branchlets obscurely quadrangular, densely tawny- or yel- 

 lowish-strigulose. Petiole 1-3 mm. Blade 4-12(-l6) x 2.5-6 mm., ovate (some- 

 times with the margins so strongly recurved as to appear oblong) with the apex 

 rounded-acute and the base broadly acute to obtuse, the 3 primaries impressed 

 above and elevated below, the secondaries obscure or obsolete; above moder- 

 ately strigulose, the slender hairs 10-15/mm. 2 and with their basal l / 3 - l / 2 adherent; 

 below densely loose-long-strigulose on the primaries, the surface moderately hir- 

 sutulous with the hairs 15-25/mm. 2 and less than 1 mm. long, the solitary glands 

 15-40/mm. 2 Flowers predominantly 5-merous, predominantly solitary on short 

 leafy lateral branchlets, closely invested by usually 4 (occasionally 6, rarely 2) 

 persistent bracts, the last branchlet leaves modified toward bracts to varying de- 

 grees; pedicel 0-10 mm. above the last branchlet leaves. Outer bracts of 6-brac- 

 teate flowers 4.5-9 x 5-7 mm., broadly ovate to orbicular with the apices broadly 

 acute to rounded; innermost 2 pairs of bracts 6-11 x 6-12 mm., orbicular to ovate- 

 orbicular with the apices rounded, outside densely sericeous-strigose with the 

 tawny to yellowish hairs 15-20/mm. 2 and to 3 mm. long. Hypanthium 5-8 x 5-8 

 mm., 0.2-0.3 mm. thick medianly, very densely sericeous-strigose with the tawny 

 hairs 6-8/mm. 2 and to 7 mm. long. Sepals 3.5*7 x 3.5-6.5 mm., ovate to oblong- 

 ovate with the apices obtuse to rounded, not or scarcely united at bases, often 

 imbricate 0.5-1 mm. Petals deep purple, 9-13 x 8-13 mm., obovate and symmet- 

 rical to slightly asymmetrical with the apices rounded or even slightly refuse, the 

 gland-tipped cilia 0.1-0.4 mm. (the terminal few 0.7-1.2 mm.). Filaments 4-6 mm.; 

 anthers 4-7 mm,; connective at anther base 0.7-1 mm., free of the anther 0.2-0.5 

 mm., the ventral lobes 0.1-0.4 ram. Style 13-25 x 0.4-0.6 mm., exserted 4-11 mm. 

 Ovary 6-7 x 3.5-4 mm., moderately to densely strigulose with gland-tipped hairs 

 on the apical 2.5-3.5 mm., the apical lobes 0.7-1.4 ram. above the locules. 



Type Collection and Locality: Bonpland s.n. (HOLOTYPE presumably in 

 Herb. Humboldt & Bonpland at P; isotype P); Ecuador, Prov. Loja, "entre Loxa 

 et Malacatos, a une elevation de 2000 metres." 



Type Photographs and Illustrations: Gleason 87-3 (holotype of B. campylan- 

 thum at K); Rhexies pi, 20 (1808) (as Rhexia conferta); Trans. Linn. Soc. 28: pi. 

 3, f. 33f (1871) (as Bracbyotum confertum); 3ot. Mag. pi. 6018 (1873), and idem, 

 Fl. Serres 20: pi. 2099 (1874) (as Bracbyotum confertum). 



Distribution: central to southern Ecuador, alt. 2000-3500 m. 



Pichincha: Cerro Corazon, Sodiro 470 (BR). Canar: between Biblian and Canar, Camp 

 E-433 (NY), Haugbt 3329 (NY, US); Pillzhum, Jameson s.n. (holotype of B. campy lanthum 

 K); near Azogues, Rose & Rose 22788 (NY, US). Azuay: near Cuenca, Jameson s.n. (BR, 

 US, W); along Rio Matadero west of Cuenca, Camp E-1994 (NY); Rio Surucuchu west of 

 Cuenca, Camp E-4203 (NY); Cumbe, Harling 852 (S); between Cum be and Nabon, Penland 

 & Summers 1094 (F, NY); Paramo de Tinajillas, Camp E-2098 (NY); between Cuenca and 

 Ona, Hitchcock 21649 (NY, US). Loja: between San Lucas and Ona, Hitchcock 21548 (NY, 

 US); Loja, Seemann 774.1 (K). 



The Jameson specimens, apart from the holotype of B. campylanthum, were 

 numbered 16, 17, and 18 by Gray, but all seem to be sheets of a single collec- 

 tion. There are no differences between B. campy lanthum and B. confertum, and, 

 in various herbaria, both Triana and Cogniaux had labeled different sheets of the 

 same collection with either one or the other of the epithets. Of the 306 examina- 

 ble flowers among the various collections, 240 were 5-merous; typical of this 

 plurality of 5-mery were such large Camp collections as E-4203 with 100 of 128 

 flowers 5-merous, E-2098 with 24 of 39, E-433 with 14 of 15, and E-1994 with 16 

 of 17. Only one sheet examined of all the collections had a majority of the flow- 



