160 MEMOIRS OF THE NEW YORK BOTANICAL GARDEN [Vol. 9, No. 2 



southwest of Cuenca, 2700-3000 m, June 29-30, 1945, Giler & Prieto {Camp E-3922); 

 vicinity of the lake in the valley of the Rio Surucuchu (a branch of the Rio Matadero), 

 18-20 km west of Cuenca, 2940-3090 m, July 16, 1945, E-4208; the eastern cordillera, 

 1-8 km north of the village of Sevilla de Oro, 2400-2700 m, July 27-Aug 12, 1945, E-4274, 

 E-4290, E-4516. E-4517, E-4518, E-4582, E-4667; ridge between El Pan and Guachapala, 

 2250-2940 m, Sept 4, 1945, E-5260. 



CA$AR: north rim of the valley of Rio de Canar, near Sus^al, April 23-25, 1945, 

 Giler (Camp E-2830); region of San Marcos, about 10 km northeast of Azogues, 2850 m, 

 Sept 25, 1944, Prieto P~79. 



CHIMBORAZO-CAflAR border (western escarpment): near Pimo, 3060-3120 m, July 9, 

 1945, E-4127. 



LOJA: Cerro Villanaco, ca 7 km west of the city of Loja, from pass to top, 2400-2850 

 m, July 28, 1944, E-234. 



Some of the specimens cited above have small leaves representative of de 

 Candolle's var. parvifolium. The variation in this character on certain individual 

 specimens, however, is so great that it is not believed worthy of taxonomic segre- 

 gation. The spikes on E-4127 are much shorter than is customary for this 

 species. This character, too, is quite variable. 



Piper nubigenum Kunth, Linnaea 13: 654. 1839. 



AZUAY: vicinity of the lake in the valley of the Rio Surucuchu (a branch of the Rio 

 Matadero), 18-20 km west of Cuenca, 2940-3090 m, July .16, 1945, E-4209. 



Piper art an the CDC. var. grand if olium Yuncker, var. nov. 



Frutex perramosus; foliis fere totis pinnatim venosis; drupa globoso-ovoidea, 

 conferte pilosa; stylo perbrevi crasso. 



Shrub, 3 m tall, with branches lax and much ramified, pubescent; leaves 6-10 

 cm wide x 14-19 cm long, pinnately nerved nearly throughout, the nerves 6-10 

 on each side; fruit globose-ovoid, densely pilose; stigmas linear, recurved, on a 

 very short, stout style. 



AZUAY: the eastern cordillera, 1-8 km north of the village of Sevilla de Oro, 2400- 

 2700 m, July 27-Aug 12, 1945, E-4481 (TYPE). 



Aside from the characters as given, in particular the comparatively large 

 leaves and densely pilose fruit, this specimen agrees very well with the Co- 

 lombian P. artanthe. 



Piper holtonii CDC. var. parvispicum Yuncker, var. nov. 

 Frutex; foliis subtus ad venas dissite villosis. 



Shrub, 1 m tall; upper internodes slender, up to 15 cm long; nerves beneath 

 and petioles loosely villous; spikes as yet young, about 2 mm thick x 5 mm long; 

 peduncle 2-3 mm long. 



CHIMBORAZO: canon of the Rio Chanchan near Huigra (mostly scrub-chaparral, with 

 a few*seepages and small swamps along the river), 1200-1350 m, May 7-14, 1945, E-3180 

 (TYPE). 



The very short spikes and villous nerves and petioles are distinctive of 

 this variety. 



Piper tuberculatum Jacq. Collect. 2: 2. pi. 211. 1788. 



GUAYAS: coastal plain, in the vicinity of Naranjito, about 36 m, June 6-7, 1945, 

 E-3545. 



Piper tuberculatum var. minus CDC in DC. Prodr. 16(1): 266. 1869. 



EL ORO: junction of Rios Ambocas and Luis, 10 km due south of Portovelo, 600-750 

 m, Oct 6, 1944, E-596. 



Piper arboreum Aublet, Fl. Guian. 1: 23. 1775. 



Junction of GUAYAS, CA5IAR, CHIMBORAZO & BOLIVAR: foothills of the eastern 

 cordillera near the village of Bucay, 300-375 m, June 8-15, 1945, E-3666 t E-3771. 



