955] 



REVISION OF THE GENUS SAGITTARIA 



197 



Perennial. Leaves emersed or submerged, the submerged leaves modified into 

 jhyllodia 4-25 x 0.4-1 cm, gradually tapering toward the tip, the emersed leaves 

 anceolate, ovate or sagittate with widely divergent lobes. Scapes simple, with 

 2-8 whorls of flowers. Bracts connate at the base, somewhat thickened, the free 

 mds triangular or elongate. Pistillate flowers with recurved, greatly inflated 

 ?edicels 3-5 cm long; sepals covering more than one-half of the receptacle at 

 naturity; petals with a purple spot at base. Stamens numerous, with glabrous 

 lilated filaments to 1.5 mm long. Mature pistillate heads 0.9-1.5 cm in diameter; 

 ichenes 2-2.5 x 1-1.4 mm, the resin duct typically absent. (FIG. 2 g.) 



Type collection: Chamisso and Schlechtendal s.rz., near the city of Concepcion, 

 Ihile (holotype not seen, presumably at Berlin). 



Distribution: Temperate Chile, from near sea level to the middle slopes of the 

 \ndes. Collections examined from the provinces of Cautin, Chiloe, Concepcion, 

 ralca, Valdivia. (FIG. 2.) 



ic. Sagittaria montevidensis subsp. calycina (Engelm.) Bogin, comb, nov. 

 Sagittaria calycina Engelm. in Torrey, Bot. Mex. Bound. 212. 1859. 

 Sagittaria calycina var. fluitans Engelm. in Torrey, Bot. Mex. Bound. 212. 1859. 

 Sagittaria calycina var. maxima Engelm. in Torrey, Bot. Mex. Bound. 212. 1859. 

 Sagittaria calycina var 6 media Engelm. in Torrey, Bot„ Mex. Bound. 212. 1859. 

 Sagittaria calycina var. grandis Engelm. in A. Gray, Man. Bot. ed. 5. 494. 1867. 

 Lophiocarpus calycinus (Engelm.) Micheli in DC. Monogr. Phan. 3: 61. 1881. 

 Lophotocarpus calycinus (Engelm.) J.G.Sm. Mem. Torrey Club 5: 25. 1894. 

 Lophotocarpus calycinus var. depauperatus Engelm. ex J.G.Sm. Missouri Bot. Gard. 



Rep. 11: 148. 1899. Nomen nudum. 

 Lophotocarpus californicus J.G.Sm. Missouri Bot. Gard. Rep. 11: 148. 1899. 

 Lophotocarpus depauperatus J.G.Sm, Missouri Bot. Gard. Rep. 11: 148. 1899. 

 Lophotocarpus fluitans (Engelm.) J.G.Sm. Missouri Bot. Gard. Rep. 11: 148. 1899. 



Annual or Perennial. Leaves emersed, floating or submerged, the emersed 

 sagittate, only rarely without basal lobes, the submerged reduced to linear phyl- 

 lodia to 45 cm long and 0.5 cm wide, the floating with ovate blades. Scape with 3-12 

 whorls of flowers, simple or branching at the lowest whorl. Bracts connate, the 

 free ends obtuse or acute, typically less than 1 cm long, occasionally to 2 cm 

 long. Pistillate flowers with thickened recurved pedicels (1-) 2.5-7.5 cm long; 

 sepals covering about one-half the receptacle at maturity; a ring of functional 

 stamens usually present. Stamens with linear pubescent filaments 1-2 mm long. 

 Mature pistillate heads 1-2.5 cm in diameter; achenes 2-2.5 x 0.9- 1.3 mm, the 

 resin duct typically present. (FIG. 8a, b.) 



Type collection: /. Hale s.n., Red River, Alexandria, Louisiana (lectotype 

 NY; isolectotypes GH, NY). Engelmann in his original description of Sagittaria 

 calycina did not cite any examples for the species. He did, however, designate 

 collections for each of the varieties proposed in the same work. The lectotype 

 chosen here is the holotype of var. maxima,, The types of the other varieties 

 proposed by Engelmann (var. fluitans and media) are treated as syntypes. 



Distribution: United States, western slopes of the Appalachians, south of the 

 Great Lakes, westward and southward to Northern Mexico and California, absent 

 from the Rocky Mountains and the Gulf states east of Louisiana. A plant of fresh 

 water (rarely saline), in ponds, lakes, sloughs, and slow moving streams. Collec- 

 tions examined from Arkansas, California, Colorado, Illinois, Indiana, Iowa, 

 Kansas, Kentucky, Louisiana, Michigan, Minnesota, Missouri, Nebraska, New 

 Mexico, Ohio, Oklahoma, South Dakota, Tennessee, Texas, West Virginia, Virginia, 

 Wisconsin; and from Coahuila, Chihuahua, and Sinaloa in Mexico. (FIG. 8.) 



