1957} 



THE BOTANY OF THE GUAYANA HIGHLAND—PART II 



375 



tubulo crassiusculo 6 mm, ligula ovato-oblonga, 3 mm lata, 7-nervata, apice 

 3-dentata. Flores disci hermaphroditi 11-12, corolla tubulosa glabra lutea, 9 mm 

 longa, tubulo 5 mm longo, dentibus linearibus, 1 mm longis. Antherae 3 mm 

 longae, basi obtusae. Rami stigmatici elongati flexuosi apice subtruncati parce 

 papillosi. Pappus stramineo-albicans. 



TYPE: VENEZUELA: Terr. Amazonas: Cerro Guanay, Cano Guaviarito, Rio 

 Manapiare, Rio Ventuari, infrequent, rocky elevations at 2,000 m alt. Subsucculent 

 vine or scandent subshrub, Bassett Maguire, Kathleen D. Phelps, Ch, B. Hitch- 

 cock & G. Budowski 31737 (NY; isotype, F). 



Senecio phelpsiae is a striking species for its thick, smooth, fleshy, nar- 

 rowly elliptical or oblong leaves. It is closely related to S. jahnii Cuatr. but 

 differs from it by the very short-petiolate and narrower leaves which have 

 another kind of nervation, by its smaller heads and flowers, and by the almost 

 complete absence of a calycle. S. cobrensis Cuatr., a related species, has 

 broader, acute, long-petiolate leaves and a different form and size of flowers. 

 S. cucullatus Klatt has broader, long-petiolate leaves and smaller heads and 

 flowers. 5. haughtii Cuatr. has richer and denser inflorescences, shorter and 

 more bracteolate pedicels and broader, long-petiolate leaves. The very deciduous 

 woolly indument on the buds and young leaves and branchlets of 5". phelpsii has 

 not been observed in the other, related species. 



Mutisieae. 



Forty-nine species of Mutisieae are presently known from Guayana. They 

 form a closely interrelated group of genera which suggest a history of closed 

 phyletic development and phytogeographically restricted distribution. Of them, 

 only one, Gongylolepis columhiana Cuatr., is extra-Guayanan (occurring in the 

 eastern Andes of Venezuela and Colombia). 



Previously to 1931, four members of this assemblage has been recognized, 

 viz., Gongylolepis benthamiana Rob. Schomb., Stifftia martiana Baker [Gongylo- 

 lepis martiana (Bak.) Cuatr. & Steyerm.], Stifftia condensata Baker [S tomato cha- 

 eta condensata (Bak.) Mag. & ^urd.], and Ouelchia conferta N. E. Brown. During 

 the period 1931-39, based on specimens collected by Tate, Blake proposed ten 

 additional names (six in Stenopadus, including S. cinereus Gleason & Blake, 

 herein transposed to Chimantaea, and S. variabilis, herein synomized under 

 Stomatochaeta condensata)^ bringing the total to 13. Now, in the past two 

 decades, contemporary exploration in Guayana has brought to light 36 additional 

 species (and 3 subspecific taxa), of which 14 have been published. Nine are 

 described herein, and 13 are to be treated in the succeeding paper. 



During the progress of botanical exploration in Guayana, there has accumu- 

 lated an impressive and significant body of evidence bearing upon fundamental 

 problems of plant phylogeny, relationships, and geography. Members of the 

 Composite Mutisieae are of special interest in this regard. The following short 

 consideration briefly summarizes arguments which support our interpretation of 

 phyletic derivation and relationship, and hierarchal arrangement of Guayana 

 mutisias. Dr. Sherwin Carlquist has examined our materials anatomically, and 

 has provided a body of data which complements our own. His studies are pre- 

 sented in papers under the titles "Anatomy of Guayana Mutisieae. I. Pollen," 

 and "Anatomy of Guayana Mutisieae. II. Floral Venation" (Mem. N. Y. Bot. 

 Gard. 9:000-000. 1957). Pollen descriptions used in this paper are furnished by 

 Dr. Carlquist. 



Subtride Gochnatinae. 



Among the gochnatinioid representatives, i.e. those with regular corollas, 



