378 



MEMOIRS OF THE NEW YORK BOTANICAL GARDEN 



[yol. 9, No. 3 



Monotypic Glossarion presents a particular problem, seeming to bridge or 

 transcent the limits of the subtribes Gochnatinae and Mutisinae. 1 ' In facies 

 the shrub recalls the stenopadoid assemblage. The ligulate corolla has five 

 equal lobes; hence, if reconstructed as a wholly gamopetalous corolla, it would 

 certainly be actinomorphic and not bilabiate. Styles are smooth as in members of 

 the Mutisinae, and the leaf base is half-clasping as in Gongylolepis. 



Key to the Subtribes and Genera of Guayana Mutisieae 



1. Corollas actinomorphic or essentially so; anthers caudately sagittate; subtribe 

 Gochnatinae. 



2. Corollas carnose, glabrous, the gamopetalous portion much exceeding the 

 tightly coiled lobes. 

 3. Corolla-tube 15 (lO)-nerved, much exceeding the tightly coiled 3 (2)-nerved 

 lobes; tails of adjacent anthers connate (but easily separable); styles 

 asperulous towards the tip, the branches 1 .0—1 .5 mm long, undif- 

 ferentiated; heads massive, terminal, solitary, many-flowered (usually 

 40-100— flowered); achenes glabrous; corona cylindric, 0.5-1.0 mm 

 high; trees. 1. Stenopadus. 



3. Corolla-tube 5-nerved, lobes 2-nerved, styles completely smooth, the 



branches compressed, dilated with marginal stigmatic lines and acute 

 appendages ca. 0.3 mm long; shrubs with slender glabrous branchlets, 

 and narrowly elliptic acute or acuminate, thin-textured leaves; shrubs; 

 confined to southern Brazil, and one species in the lower Rio Negro ba- 

 sin. (Stif/tia.) 

 2. Corollas firm or submembranous , but not carnose; the gamopetalous portion 



equaling or considerably shorter than the erect or merely recurved 2-nerved 



lobes. 



4. Heads solitary and terminal; corollas hirsute within the collar. 



5. Heads relatively large, 20-50-flowered; corolla-tubes 5-nerved, the lobes 

 densely barbate-pilose at the tips (this pubescence largely deciduous 

 at anthesis), erect or sometimes somewhat recurved; anther-tails all 

 free; styles asperulous upwards, the branches undifferentiated, 0.7-1.0 

 mm long; achenes pubescent or glabrous, 6-9 mm long; corona 1.0-1.5 

 mm long, deeply 5-lobed; virgate or little- branched ^depressed, sub- 

 succulent shrubs. 2. Chimantaecu 

 5. Heads relatively small, 10-20 (30)-flowered; corolla-tube 10-nerved, the 

 lobes glabrous, stiffly erect, 2-nerved; tails of adjacent anthers con- 

 nate (but easily separable); styles asperulous upwards, the branches 

 obtuse, ca. 0.5 mm long; achenes 6 mm or less long, glabrous, corona 

 entire, 0.5 mm or less long; brittle, much-branched shrubs, the pubes- 

 cence malpighioid. 3. Stomatochaeta. 

 4. Heads uniflorous, in axillary cymes; corollas glabrous within; tube 5- 

 nerved, the lobes recurved, 2-nerved; style smooth, glabrous, the branches 

 1.2-1.75 mm long, 3-lobed; achenes glabrous, 4-6 mm long; corona 

 cylindric, 0.3-0.5 mm long; upright, branched shrubs. 4. Quelchia. 

 1. Corailas ligulate, the ligule 5-oblong-dentate, 7-10-nerved; heads 12-16- 



flowered; anthers caudately sagittate; shrubs; subtribe Mutisinae. 5. Glossarion. 

 1. Corollas bilabiate; subtribe Mutisinae. 



6. Heads 8-« flowered, massive, terminal, solitary or pedunculate, the pedun- 

 cles sometimes branched. 

 7. Leaves pinninerved; anthers caudately sagittate, the adjacent tails con- 

 nate; achenes glabrous or essentially so; phyllaries broadly obtuse, 

 more or less coriaceous. 6. Gongylolepis. 



7. Leaves 1-nerved; anthers caudately sagittate, the tails free; achenes 

 obviously pubescent; phyllaries acute or acuminate, more or less per- 

 gameneous or papyraceous. . 7. Duidaecu 

 6. Heads small, 2-5- flowered, the inflorescences axillary, in 3-7-headed cymes. 



"We have attempted to accomodate our material to the organization of subtribes as 

 presented by Hoffmann. It would seem, however, that fresh evaluation of the entire 



Mutisieae is now needed. 



