1957] 



THE BOTANY OF THE GUAYANA HIGHLAND— PART II 



379 



8. Phyllaries glabrous, the margins ciliolate; corolla funnel-formed, glabrous; 

 caudal appendage of anther sagittate, the tails of adjacent anthers con- 

 nate; style-branch tips ca. 1 mm long, rounded, undifferentiated; achene 

 6-7 mm long, 5-nerved; pappus 3-seriate; leaves oblanceolate. 8. Neblinaea, 



8. Phyllaries strongly externally bearded; corolla goblet-shaped, puberulent; 

 caudal appendage of anther linear, simple, not sagittate, not connate; 

 style-branch tips obtuse, dorsally unilobate; achene 5-6 mm long, 10- 

 nerved; pappus 5-seriate; leaves elliptic. 9. Achnopogoru 



1. Stenopadus Blake, Bull. Torrey Club 58: 489. 1931, descr. emend. 



In 1953* we recognized six species in Stenopadus proper {Eustenopadus 

 Blake), all endemic to the Guayana Highland. Now, as a result of further explo- 

 ration, eight additional species have been collected (two of them to be described 

 in the succeeding paper) in Venezuelan Guayana, and one, 5". colombianus Cuatr. 

 & Steyerm., has been described from Colombian Guayana. 



Heads massive, terminal, solitary (in three species sometimes 2-4), homoga- 

 mous, multiflowered [(15) 40-100]; receptacle glabrous, alveolate, strongly or 

 sometimes not at all paleaceous; corolla actinomorphic, carnose, essentially 

 infundibul iform and differentiated into a cylindric tube, an expanded limb, and 

 tightly coiled, linear-lanceolate lobes, the tubular portion much exceeding the 

 lobes, characteristically 3 (sometimes only 2) distinct veins extending from the 

 base of the corollas into the lobes; anthers caudate-sagittate, the adjacent tails 

 connate; filaments attached either in the sinus of the lobes, at the union of the 

 tube and the limb, or below this union; pollen grains yellowish, tricolpate, the 

 furrows long-pointed; grains prolate to subprolate (prolate-spheroidal in 5". 

 campestris), spinulose or psilate; polar axis 55-65 fi; achenes prismatic, basi- 

 cally 10-nerved, glabrous; annulus mostly lacking; corona cylindric; pappus- 

 setae mostly subpaleaceous, flattened and connate into a ring at the base. 

 Trees. 



TYPE species: Stenopadus talaumifolius Blake. 



Key to the Sections and Species of Stenopadus 



1. Leaves coriaceous, the lateral veins prominulous but the blades not reticulate- 

 veined. 



2. Filaments attached within the corolla at the union of tube and limb, hence at 

 or below the middle of the gamopetalous portion and not at the sinus; 

 receptacle strongly paleaceous; achene obviously 10-nerved; annulus 

 lacking; setae subpaleaceous, 5-8-seriate, subconnate at the base. 

 3. Corolla infundibuliform, (18) 20-40 mm long, the lobes tightly coiled; 

 filaments attached at the middle; sect. Kunhardtia. 

 4. Involucre broadly campanulate, 60-1 00- flowered; leaf-blades broadly 

 obovate or obovate-oblanceolate, usually 8-25 cm long, 4-8 cm broad. 

 5. Involucre substipitate; heads 60-70-f lowered; corollas ca. 40 mm long; 

 anthers ca. 15 mm long; leaves usually 10-25 cm long, 6-8 cm 

 broad. 1. Stenopadus kunhardtiu 



5. Involucre sessile; heads 80- 100-f lowered; corollas ca. 30 mm long; 



anthers ca. 10 mm long; leaves usually 8-12 cm long, 3.5-6.0 cm 

 broad. 2. Stenopadus huachamacari. 



4. Involucre obconic or hypo crater iform, 2 5-3 0-f lowered; leaves obovate 

 or oblanceolate, usually 6-12 cm long, 2-5 cm broad. 



6. Phyllaries obtuse; involucre hypocrateriform, strongly stipitate; heads 



40-50-flowered; corollas 18-20 mm long; leaf-blades narrowly 

 oblanceolate, 7-12 cm long, 3-5 cm broad; petioles 15-22 mm long. 



3. Stenopadus stipitatus. 



*Maguire, B., Cowan, R. S. & Wurdack. J. J. The botany of the Guayana Highland. 

 Mem. N. Y. Bot. Gard. 8: 153. 1953. 



