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MEMOIRS OF THE NEW YORK BOTANICAL GARDEN 



[Vt>L 9, No. 3 



the achene top, enter the achene (fig. 109 C). Five other bundles, terminating 

 upwardly at the achene top, are also present as exterior bundles in the achene. 

 The style (fig. 109 B) contains the usual dorsiventral pair of bundles. At lower 

 levels in the style, a pair of lateral bundles also is found, however. These four 

 style bundles enter the achene (fig. 109 C); the dorsiventral pair unites with 

 exterior bundles a short distance below the summit of the achene (fig. 109 E). The 

 lateral pair continue down the achene as interior bundles (fig. 109 E, F). Near 

 the base of the achene, the interior bundles fuse with the ovule trace; this group- 

 ing joins at the base of the achene a grouping formed from the exterior bundles. 

 A single vascular strand connects the achene vascular tissue with that of the 

 receptacle. Specimens examined of other species of Quelchia, 0. conferta, Q. 

 eriocaulon, and Q. x grandifolia, were found to have a floral venation identical to 

 that seen in Q. cardonae. 



Neblinaea. 



A number of similarities may be found between those genera of the present 

 study having actinomorphic corollas (Gochnatinae) and those possessing bilabiate 

 corollas (Gerberinae). Flowers of Neblinaea (fig. 110) are exemplary in this 

 respect. Corollas of the only species, N. promontorium (fig. 110A), show lateral 

 bundles in the two free lobes. It is of considerable interest to find that the 

 lateral bundles of the three united lobes are not united as they are in bilabiate 

 flowers of more advanced Mutisieae, such as are described below. The two pairs 

 of adjacent "laterals" of this portion of the corolla fuse in the basal portion of 

 the corolla, or in the achene top. Beneath the three sinuses beside the two lobes 

 which are not united, however, the laterals fuse immediately below the sinuses. 

 In addition to the full representation of lateral bundles in this corolla, median 

 bundles are occasionally found either in the strap-shaped portion of the corolla or 

 in the lobes which are separate. These median bundles terminate freely at both 

 ends and extend only a short distance into the corolla tube. Stamen traces unite 

 with corolla bundles in the top of the achene. As in the three preceding genera, 

 ten bundles are present in the achene (fig. 110 C), five of these being additional 

 to those connected to corolla bundles. As in Quelchia, four bundles are present 

 in the style (fig. 110 B); the two lateral style bundles fuse with the dorsiventral 

 pair at various points above the base. The achene structure is also identical to 

 that of Quelchia in that the dorsiventral style bundles fuse with external achene 

 wall bundles in the achene top (fig. 110 E), while the lateral style bundles con- 

 tinue into the achene as interior bundles accompanying the bands of stigmatoid 

 tissue. The fusion of all bundles into a single strand of vascular tissue at the 

 base of the achene is also like the condition found in Quelchia. The similarities 

 of floral venation between Quelchia and Neblinaea are not cited here to suggest a 

 close relationship between these genera. Rather, simplifications (as compared to 

 genera described above) such as they show may occur independently. 



Gongylolepis 



The genus Gongylolepis is interesting to compare with Neblinaea in that 

 although the flowers of most species are much larger than those of Neblinaea, 

 their flowers do not have more complex venation; in fact, their pattern is often 

 simpler than that just described. In Gongylolepis bracteata (fig. Ill) the con- 

 dition might be interpreted as basic for the genus, though it is not the most 

 typical. In the corollas (fig. Ill A) of this species lateral veins are present, 

 fusing at the lobe-tips both in the two separate lobes and the three united lobes. 

 The latter portion of the corolla exhibits a very interesting condition (mentioned 



