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MEMOIRS OF THE NEW YORK BOTANICAL GARDEN 



[Vol. 9, No. 3 



corolla bundles in the achene top. In addition to the corolline bundles, five more 

 exterior bundles are found in the achene wall (fig. Ill C, F). The style of 

 Gongylolepis bracteata (fig. Ill B) has four bundles at the base (fig. Ill D). In 

 its upper portions, however, the bundles branch and anastamose, forming a more 

 complex appearance; these groups of bundles are arranged on adaxial and abaxial 

 sides of the style, and pass into the stigmatic branches. In the achene (fig. 

 Ill C), the dorsiventral style bundles unite with exterior achene wall bundles on 

 opposite sides of the achene. The lateral style bundles continue down the achene 

 as interior bundles (see also fig. Ill F). Near the base of the achene these 

 bundles unite with the ovule trace (fig. Ill G) to form a cylindrical plexus. Further 

 down, this joins with the exterior bundles to form a single strand which connects 

 with the receptacle. The ovule trace was found to be dual along most of its 

 length. 



The drawing of Gongylolepis pedunculata (fig. 112) shows the only species 

 of this genus studied in which a more complex corolla venation was found. In 

 certain features it is like the condition in G. bracteata. A median vein, for ex- 

 ample, is present for most of the length of the strap-like portion of the corolla. 

 As in G. bracteata, two prominent veins from "laterals'' beside the central two 

 veins which run the length of this structure. These branch at the same level as 

 the veins subtending sinuses beside the separate lobes. In addition, other veins 

 branch off from the five main corolla-tube veins at this level. The shorter, freely 

 terminating branches, unlike those of Stenopadus kunbardtii or Stomatochaeta 

 crassijolia, are located inside, rather than outside, the true laterals. The style of 

 G. pedunculata (fig. 112B) is simpler than that of G. bracteata in that only four 

 bundles, all of which terminate freely, are found. The achene venation is like that 

 of G. bracteata. The greater complexity of the corolla venation in G. bracteata 

 may be correlated with the fact that this species has the largest flowers of the 

 species of Gongylolepis studied. 



The smallest- flowered species, G. fruticosa, (fig. 113), has been included for 

 comparison. In the corolla of this species (fig. 113 A), the venation has reached 

 the ultimate simplicity found in most zygomorphic corollas of Mutisieae. The 

 laterals fuse directly beneath the sinuses, and no corolla veins are found in 

 addition to the fused laterals. In the achene (fig. 113 C) the five corolline bundles 

 are augmented, as in G. bracteata, by five more bundles, which terminate in the 

 top of the achene. In the style of the flower of G. fruticosa studied (fig. 113 B), 

 four bundles were present, though only three of these extended to the style 

 base. In the achene (fig. 113 C), these were found to join achene wall bundles at 

 a high fevel of the achene. Thus only exterior bundles were present in most of the 

 length of the achene. A simple ovule trace is present; all the bundles of the 

 achene unite at the achene base into a single strand of vascular tissue. 



Despite the fact that their corollas are larger than those of G. fruticosa, the 

 same corolla venation was seen in G. huachamacari, G. martianus, and G. paruana. 

 The styles were all found to have four bundles; a pair of lateral interior bundles 

 were usually not observed in achenes of th£se species, all the stylar bundles 

 uniting with achene wall bundles at the summit of the achene. The ovule trace 

 was often seen to be dual, at -least in its upper portion. 



Explanation of Figures 112-113 



FIG. 112. Gongylolepis pedunculata {Cowan & Wurdack 31131), portions of flower 

 drawn to show venation. A, corolla; B, style. X 4.5. FIG. 113. Gongylolepis fruticosa 

 {Steyermark & Wurdack 1136), portions of flower drawn to show venation: A, corolla; B, 

 style; C, achene. A, B, x 4.5; C, X 9. 



