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Diphyllostomatidae: Diphyllostoma 



Lucanidae: Aegus, Aesalus, Ceruchus, Chiasognathus, Dörens, Figulus, Lamprima, 

 Neolucanus, Nicagus, Nigidius, Penichrolucanus, Platycerus, Prosopocoilus, Sinoden- 

 dron, Syndesus 



Glaphyridae: Amphicoma, Lichnanthe 



Trogidae: Trox, Omorgus, Polynoncus 



Bolboceratidae: Athyreus, Australobolbus, Blackbolbus, Blackbiirninm, Bolbaineus, 

 Bolbcipium, Bolboceras, Bolbocerastes, Bolbocerosoma,bolbocerosum, Bolbochromus, 

 Bolbogoniwn, Bolbohamatum, Bolbelasmus, Bolboleaus, Bolborhachium, Bolborhinum, 

 Bolborhombus, Bradycinetulus, Elephastomus, Eucanthus, Gilletinus, Neoathyreus, 

 Pereirabolbus, Stenaspidius 



Pleocomidae: Pleocoma 



Geotrupidae: Anoplotrupes, Ceratophyus, Ceratotrupes, Chromogeotrupes, Cnemotrupes, 

 Enoplotrupes, Epigeotrupes, Frickius, Geohowdenins, Geotrupes, Haplogeotrupes, 

 Megatrupes, Mycotrupes, Odontotrupes, Onthotrupes, Phelotrupes, Sericotrupes, 

 Thorectes, Typhoeus 



Hybosoridae: Anaides, Araeotanopus, Brenskea, Chaetodus, Dalmothoracodes, 



Hapalonychus, Hybochaetodus, Hybosorus, Liparochrus, Microphaeochroops, 



Phaeochridius, Phaeochroops, Phaeochrous, Triclwps 

 Ceratocanthidae: Astaenomoechus, Ceratocanthus, Cloeotus, Cyphopisthes, Eubrittoniella, 



Eusphaeropeltis, Madrasostes, Perignamptus, Philharmostes, Pterorthochaetes, 



Synannostes 



Ochodaeidae: Chaetocanthus, Ochodaeus, Pseudochodaeus, Synochodaeus 

 Scarabaeidae: Acognatha, Acoma, Aegialia, Agamopus, Alaberoides, Allokotarsa, 

 Amphimallon, Anachalcos, Anatochüus, Anisonyx, Anomala, Anomaluera, Aphodins, 

 Aphonides, Apogonia, Archophilenrns, Asthenopholis, Ataenius, Aulonocnemis, Bolax, 

 Brachymacroma, Callirhinus, Camenta, Camentoserica, Campilipns, Campsiura, 

 Catharsins, Canthidium, Canthon, Carhvrightia, Chironitis, Chlorocala, Chnaunanthus, 

 Circellium, Coenochilus, Colobopterns, Comythovalgits, Copris, Coprophanaens, 

 Coptorhina, Cotinns, Cyclocephala, Cyclomera, Cymophoms, Cyphonistes, 

 Cyptochirns, Cyrioperta, Deltochilnm, Deltorrhinum, Diastictns, Dichelonyx, Dichelns, 

 Dichotomins, Dinacoma, Diplognatha, Diplotaxis, Drepanocanthns, Drepanocems, 

 Drepanopodns, Dynastes, Dyscinetns, Eriesthis, Enoniticelhis, Eitparia, Eurysternns, 

 Entheola, Garetta, Geniates, Gemichns, Gnorimella, Golofa, Gymnoloma, 

 Gymnoplennis, Heliocopris, Heteronychns, Hoplia, Hybaloides, Hybalns, Hyboscherna, 

 Hypselogenia, Kheper, Lampea, Lepidota, Lepithrix, Leptohoplia, Lencothyrens, 

 Liatongus, Macrodactylns, Melinesthes, Milichns, Neoserica, Nyassinns, Olbaberoides, 

 Oncerns, Oniticellns, Onitis, Onthophagns, Oplostomus, Orizabus, Orphnidus, Orphans. 

 Osmoderma, Oxygrylins, Oxysternon, Pachycnema, Paracotalpa, Parathyce, Pedaria, 

 Pedaridinm, Pelidnota, Peritrichia, Phacosoma, Phalogogonia, Phalops, Phanaeus, 

 Philenrns, Philoscaptns, Phobetns, Phyllopertha, Phyllophaga, Popillia, Proagoderus, 

 Psendachloa, Psendataenius, Psendorphnus, Pycnoschema, Raceloma, Rhinocoeta, 

 Rhyssemns, Sarophorus, Scarabaens, Scatimns, Sceliages, Scelophysa, Schizonycha, 



