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Figs. 9- 15: 9: Dorsolateral view of the third axillary of Glaresis walzlae (Glaresidae) showing original 

 band sclerites and detached AXCu fragment of 3Ax. - 10: Dorsal view of the first basal plate of 

 Glaresis walzlae (Glaresidae) showing HP, PC+C. 1 Ax-BScA articulation point, BScA, ScA and BRP 

 - 11: Dorsal view of the first basal plate (excluding BRP) of Scarabaeinae (Scarabaeidae) showing 

 the humeral plate (HP); the proximal and distal sections of BSca, and ScA bulge. - 12: Anterior view 

 of the humeral plate of Scarabaeinae (Scarabaeidae) showing the proximal, dorsal, ventral and distal 

 margins. - 13: Second basal plate (including BRP) of Scarabaeoidea: a, Glaresis walzlae (Glaresidae) 

 showing BRP, proximal and distal arches of BRP, embayment (open), RP, brp, brp projection, BMA. 

 BMP, BMP-BCuA brace and BCuA; b. Bolboceratidae showing BRP, embayment (closed), RP, BMA, 

 the proximal and distal arches of BMA, BMP, MP BMP-CuA brace, BCuA. BCuP, distal embayment 

 and CuA. - 14: Dorso-lateral view of the basalare of Bolboceratidae (Geotrupidae) showing the HP 

 lobe and BScP lobe. - 15: Dorso-lateral view of BScP of Bolboceratidae, showing BScP and ScP. 

 Not to scale. Originals Browne (1993). 



