17 



The ventral wall of the Ive-pouch is almost completely membranous, and it is at the same 

 time the left-anterior part of the dorsal wall of a large ventral lobe (via in fig.321e,g,i). 

 The ejaculatory duct (D in fig. 320, 32 Ig) opens into this wall. In some species this wall 

 contains a small sclerite L5 (fig.321e,g). The ventral wall of the vla-lobe is part of the 

 ventral wall of the whole left complex and is largely occupied by the right-ventral L4- 

 sclerite (fig.321g,i). 



All Blattaria, but not Mantodea, have a large hook-like evagination from the left ventral 

 wall of the left complex (hla in fig.321i). The hla-hook is largely occupied by sclerite 

 L3, but a more or less extensive basal part is membranous (30 in fig.321i). This membrane 

 can be introverted, which procedure results in a more or less deep retraction of the hook 

 into the left complex. 



Right phallomere 



The anteriormost ventral wall is occupied by the plate-like sclerite R3 (fig.321f,h). Along 

 the lateral and anterior margins of R3 (parts of the basal line Bl) the sclerotisation of R3 

 extends somewhat into the wall of the genital pouch, and these margins of sclerite R3 

 form a groove- (as seen from externally) or fold-like (as seen from internally) apodeme 

 age (fig.321f,h). Like in the swe-apodeme, parts of this age-apodeme can be filled in by 

 the cuticle becoming thickened, and the respective parts of age are beam-like. 

 Behind the central part of R3 the ventral wall is extensively invaginated dorsad and 

 anteriad (cbe in fig.321f,h), and this invagination is partly sclerotised in its dorsal wall 

 (anterior part of sclerite Rl in fig. 32 If). Blattaria, but not Mantodea, have a sclerite R2 

 left- ventral to the cbe-invagination (fig.321f,h), which articulates with the left posterior 

 end of R3 (articulation A7 in fig. 32 If) and with the left end of Rl (articulation A6 in 

 fig. 32 If). Sclerite R2 and the posterior margin of the anterior part of Rl often form tooth- 

 or ridge-like cuticular evaginations (on Rl: pva in fig.321h). 



The part of the right phallomere posterodorsal to the cbe-invagination is composed of a 

 large dorsal lobe (fda in fig. 32 If) and a ventral tooth (pia in fig.321h, which is in most 

 species much smaller than the fda-lobe). The fda-lobe and the pia-tooth are confluent 

 along the right edge of the right phallomere, and they diverge to the left. The dorsal wall 

 of the fda-lobe - and often parts of its ventral wall, too - as well as the dorsal and ventral 

 walls of the pia-tooth are occupied by the posterior part of sclerite Rl. The sclerites Rl 

 and R3 articulate with each other at the anterior right edge of the right phallomere 

 (articulation A3 in fig.321f,h). In the anteriormost dorsal wall of the fda-lobe, part of the 

 cuticle is invaginated to form a hollow, long and narrow, membranous tendon (tre in e.g. 

 fig.74; not shown in fig.321), which is present in some Blattaria only. 



Discussion of the basic division of the phallomere complex 



I propose this division of the phallomere complex into a left complex and a right 

 phallomere. However, earlier suggestions for a basic division differ from this hypothesis: 

 Snodgrass (1936, 1937) divides the phallomere complex of Blattinae into a ventral, a right, 

 and a left phallomere. Beier (1970) follows Snodgrass regarding Blattaria as well as 

 Mantodea, and he terms these main divisions hypophallus, right epiphallus, and left 



