24 



(e.g. processes or pouches) on or in which these sclerites are situated. For example, lamina 

 ventrale (= Iv) designates the sclerite I call L2 and, at the same time, the pouch Ive, which 

 contains sclerite L2 (fig.321g). This ambiguity makes LaGreca's terminology rather 

 impractical. The other terminologies put forward (e.g. Beier 1968) are not very handy 

 because of their long terms, and they are by far not detailed enough for my purposes. 



Blattaria 



McKittrick (1964) has developed a new, very simple and handy terminology, which has 

 been adopted by most of the subsequent workers. Mizukubo & Hirashima (1987) also 

 employ it but propose some changes. In both terminologies, the names for sclerites are 

 short sequences of letters and numbers, each position containing certain information. Some 

 of these terms have already been applied in the description of the phallomeres in 3.1., e.g. 

 L2, R3. 



McKittrick regards the phallomere complex of Cryptocercus punctulatus as the most 

 primitive and takes it as the reference type for her terminology. She adopts the tripartition 

 of Snodgrass (1936) into left, right and ventral phallomeres, and according to this major 

 division McKittrick basically distinguishes left, right and ventral sclerites, which get L, 

 R, or V, respectively, in the first position of their names. Then, on the left and on the 

 right phallomere, the sclerites are numbered separately. The ventral phallomere has only 

 one sclerite. In this way seven main sclerites are distinguished (LI, L2, L3, V, Rl, R2, 

 R3). No assumptions concerning side-homologies are intended in this terminology. What 

 McKittrick - starting fom this situation in Cryptocercus - regards as a product of a 

 secondary "subdivision" or as a special region or "elaboration" of a main sclerite is 

 expressed by the addition of one or two small letters (d = dorsal, v = ventral, I = lateral, 

 m = median, vm = ventromedian). Sclerites of certain species regarded as completely new 

 elements not present in Cryptocercus are given the next free number of the respective 

 phallomere. This terminology is very handy and clear and contains a lot of information. 

 Mizukubo & Hirashima state side-homologies for the elements of the left and of the 

 right half of the phallomere complex and integrate these assumptions in their terminology. 

 For that purpose, they modify the terminology of McKittrick in two ways: 



(1) According to the assumption of a plane of symmetry, they basically distinguish right 

 and left elements (R or L in first position); then both R and L are grouped into dorsal 

 and ventral elements (D or V in second position). The left-dorsal elements LD and the 

 left- ventral elements LV compose the left phallomere. The right- ventral elements RV 

 correspond to the ventral phallomere (vla-lobe in my terminology). The right-dorsal 

 elements RD correspond to the right phallomere. Thus, the basic division into LD, LV, 

 RV, and RD essentially conforms with the division of the phallomere-complex proposed 

 by Quadri (1940). 



(2) As regards the numbers and small letters, Mizukubo & Hirashima adopt the 

 terminology of McKittrick, but changes are made in order to get side-homologous elements 

 provided with the same names - except for R or L in the first position. These changes 

 are, compared with McKittrick, not very extensive. 



