44 



posteriorly onto two processes (fig. 20, 27): the left pda-process and the right vla-lobe. 

 L4A and L4B articulate on the left edge of the left complex (Al in fig. 20, 24). L4A bears, 

 anterior to Al, a small keel-like apodeme (swe in fig. 23, 24; cross-section in fig. 25). 

 Like Sphodromantis, Metallyticus has a dorsal pouch pne and a ventral pouch Ive, which 

 contain the sclerites LI and L2 (fig. 23-25). Within the pne-pouch (fig. 24, 25) LI occupies 

 the ventral wall and parts of the dorsal wall. In the ventral wall LI is partly divided by 

 a stripe of membrane (2 in fig. 25). A phallomere-gland, whose opening were to be expected 

 within the pne-pouch, was not found. The ventral pouch Ive (fig.26) deepens strongly 

 anteriad. Its dorsal wall is occupied by the ribbon-shaped L2-sclerite. The edge between 

 the pouches pne and Ive (1 in fig.25, 26) has its anterior starting point in the ventral wall 

 of the pne-pouch. The anteriormost part of edge 1 bulges to the right (afa in fig.25, 26). 

 Most posteriorly edge 1 bears a somewhat hook-like process paa, whose ventral wall is 

 sclerotised by a posterior extension of L2 (fig. 20, 25, 26). Another sclerotised process loa 

 (fig. 23-25) arises from the posterior ventral wall of the pne-pouch (fig.25). Anterior to 

 the loa-process, in the area of the bulge afa, LI bends around the edge 1 into the dorsal 

 wall of the Ive-pouch, where it has a hinge-like articulation with the right margin of L2 

 (A2 in fig.26). 



The membranous ventral wall of the Ive-pouch is at the same time the left part of the 

 dorsal wall of the ventral lobe via (= ventral phallomere; fig.27). The ejaculatory duct (D 

 in fig.26, 27) opens into the anterior ventral wall of the vla-lobe. Dorsal to the genital 

 opening there are two membranous lobes (goa in fig.25, 26). Posterior to the genital 

 opening the dorsal vla-wall contains a small sclerite L5, with some small but distinct folds 

 to the left of it. The ventral vla-wall is completely sclerotised by a part of L4A (fig.20). 



Right phallomere 



Sclerite R3 occupies the anterior ventral wall (fig.20, 24). The left end of R3 has a short 

 extension to the posterior (fig.20, 26). The anterior and right margins of R3 form a groove- 

 like apodeme age, which deepens abruptly in its left part but does not curve like in 

 Sphodromantis (fig.20, 24; cross-section in fig.21). Next to its right-posterior end the age- 

 apodeme bears a small keel-like apodeme (3 in fig.20, 24). The right posterior end of R3 

 articulates (A3 in fig.21, 24, 26) with the posteroventral sclerite RIC. Sclerite RID is 

 situated to the left of RIC and posterior to R3; it is in close contact with RIC and bears 

 a dental ridge (pva in fig.20, 25, 26). The ventral wall of the right phallomere posterior 

 to R3 and RID is extensively invaginated dorsad and anteriad (cbe in fig.21, 24). Sclerite 

 RIC extends from articulation A3 posteriad onto a large ventral tooth (pia in fig.20, 26), 

 occupying mainly its dorsal wall. Like in Sphodromantis, the area of articulation A3 and 

 the adjacent part of RIC are somewhat groove-like - a posterior extension of the age- 

 apodeme on R3. The posterior part of the right phallomere is composed of a large dorsal 

 lobe (fda in fig. 23, 24) and of the ventral pia-tooth. The dorsal wall of the fda-lobe is 



Figs. 24-27: Metallyticus violaceus (Mantodea, Metallyticidae) - Left complex and right phallomere 

 in dorsal view; with successive removal of their parts (mainly of dorsal ones); fig.27: all parts of 

 right phallomere removed. - Scale: 1mm. 



