187 



(as in Eurycotis, see below). The musculature of these species (not studied in Archiblatta) 

 confirms the assumed homologies for LI and pne: Like in e.g. Mantoida or Cryptocercus, 

 there is a stout muscle from Lla to L4-sclerotisations (compare fig. 53: L4C) in the left 

 edge of the left complex (12) and another one to L2 (compare fig.55) in the dorsal wall 

 of the Ive-pouch (13). Muscle 11 is missing. A derived feature peculiar to Blattinae (and 

 Eurycotis, fig. 70) is the shift of the left insertion of muscle b4b to the anterior summit of 

 the pne-pouch (discussion in 6.7.1.). 



In Eurycotis (fig. 65-67) the characteristics of Lla, Lll, Llm, and A2 (fig.323e) and the 

 position of the phallomere-gland opening (P and edge 6 in fig. 54, 55, 67, 68) are quite 

 the same as in Archiblatta. The pouch-shape of pne, however, is by far less distinct. The 

 processes posterior to LI could be dca (as labelled in fig. 66, 67) or loa (the right one?). 

 The insertion of 12 (fig.70) is still on the left part of the pne-pouch but has shifted from 

 LI to the adjacent membrane. (The position of the 12-insertion on L4 is the same as e.g. 

 in Mantoida: discussion in 6.3.1.). Like in Blattinae, muscle 11 is missing. Muscle 13 from 

 LI to L2 is represented by three bundles (13a,b,c in fig.71), which together occupy the 

 same insertion area as the 13 of Blattinae, and an apomorphic tripartition can be assumed. 

 Muscle b4b inserts, like in the Blattinae, anteriorly on the pne-pouch (fig.70). The origin 

 and homology of the sclerites L6A and L6B (fig. 66, 322g) only found in Eurycotis is 

 questionable: new sclerites or derivatives of LI? Homology with sclerite L8 of Ergaula, 

 Polyphaga (fig. 117, 3221,m), and Lamproblatta (fig. 177, 322k) is unlikely (different 

 muscle insertions); homology with L9 of Ergaula (fig. 105) is also not very probable. 



Lamproblatta 



Like in the other species, LI lies in the dorsal wall of the left complex, its anterior part 

 Lla (fig. 323k) is inside a deep pouch (pne in fig. 177), and its right part articulates with 

 L2 (A2 in fig. 178). Furthermore, LI and pne can be identified by the characteristic muscle 

 connections with the area of L4 in the left edge of the left complex (12 in fig. 184; the 

 L4-sclerotisations are highly modified, discussion in 6.3.4.) as well as with L2 (13 in 

 fig. 187; the insertion on L2 is far posteriorly). Like in Polyphaga, Ergaula, and Crypto- 

 cercus, the 12-insertion on LI has shifted far anteriad. Muscle U is missing. 

 As compared with other Blattaria and Mantodea, LI and pne have shifted right-anteriad. 

 Most of the anterior part of LI (Lla in fig. 323k) is level, but, in contrast to Blattinae, 

 Eurycotis, and Tryonicus, there is a reminiscence of the hood-shape since the anteriormost 

 part of LI bends into the dorsal wall of pne (fig. 177, 178). This dorsal part of Lla may 

 even be regarded as a vestige of an anterior plateau which has been inclined posteriad. 

 Sclerite arms (regions Lll and Llm) are not distinct. The part of LI containing articulation 

 A2 can be designated as the vestigial Llm-region (fig. 323k; that A2 in fig. 178 really is 

 A2 is shown in 6.2.4.). The demarcation of Lll in fig. 323k is tentative. For the process 

 dca (fig. 177) the homology with the dca (or loa, fig. 54?) of the other species is 

 questionable. Region Llr is missing (no sclerite ring). The phallomere-gland opens, hke 

 in Archiblatta or Ergaula, into the membrane extending ventrad from the posterior margin 

 of LI (P in fig. 178); however, parts of L2 and L4 (with the processes paa and pda, 

 fig. 178) have shifted into the interspace between LI and the opening (compare in 6.6.4.). 



